Concept

Voronoi cells — where it appears

4 essays name this object, across one field. What follows is each of them, and the objects they name alongside it.
5 sides13420%6 sides45869%7 sides7311%665 bounded cellsmean 5.908 sidessix is forced, not chosen

Why the average cell has six sides

Not because hexagons are efficient. Because Euler's formula leaves a tiling no choice — count the edges two ways and the mean comes out at six, whatever the cells would prefer. The efficiency argument is a different claim about a different thing.

tissue · sixsides
00.2500.5000.7501120130140150divergence angle (°)closest pair, as a fraction of the mean spacing (higher is better)137.508°400 points per anglethree criteria, three winners

Packing, measured four ways

The claim is that the golden angle packs best. It is measurable, and the measurement gives three different winners on three criteria — all near 137.5° and none of them it. That does not make the claim wrong; it makes the usual statement of it wrong.

tissue · packing
00.500155.5066.507sides of the cellarea, as a multiple of the mean cell areaLewis269 interior cellsslope 0.014 against 0.25

Lewis's law wants disorder

Cell area rises linearly with side count — measured on cucumber epidermis in 1928 and quoted ever since as a property of packed tissue. It holds beautifully on a random point set, with a fitted constant of 1.64 against Lewis's 2. On a phyllotactic head it does not hold at all: the slope is 0.009, and area and side count are almost independent.

tissue · lewis
Lewis slope — golden head0.009the law says 0.25Lewis slope — random set0.231the law says 0.25Aboav a — golden head1.177the law says 1.2Aboav a — random set0.593the law says 1.2filled where the tiling obeys the law it is being judged by900 points in each tilingLewis explains 32% of the area spread at best

Two laws that want opposite tissue

Lewis's law and Aboav's relation are quoted side by side as properties of cellular tissue. Measured on the same two tilings they point opposite ways — the ordered head satisfies Aboav's with the textbook value of 1.18 and fails Lewis's completely; the random set does exactly the reverse.

tissue · cell laws

Named alongside it

The objects these essays reach for when they reach for this one.

Divergence angleLewis's lawOrder and disorderAboav–WeaireAsymptotic claimCriterion dependenceCylinderEpitheliumEuler's formulaExplained varianceHexagonJoint distribution

All concepts