Phyllotaxis
the spirals are counted, not admired
Theme

The thread: The angle is an output

The golden angle is where a rule settles, not a number the rule contains. Sweeping the one parameter of that rule gives a diagram with a golden branch, a transition and a two-whorl regime — and the famous constant is one branch of it.
The rule, 26 steps in, at a growth of 0.40. The next primordium goes where the repulsion is least — the marked minimum at 216°. Nothing in the rule refers to any particular angle. Where the angle comes from

The angle is an output

137.5° is not a constant of nature. It is where a rule settles — a rule that places each new element as far as it can from the ones already there, contains no reference to the golden ratio, and reaches the same answer from starting angles a hundred and sixty degrees apart.

 The spiral counts four different divergence angles produce. Fibonacci counts come from one angle. The Lucas angle — which the same dynamical model reaches on a different branch — gives 47 and 76, and neither number is a Fibonacci number. The claims, measured

Fibonacci is a branch, not a law

Fibonacci counts come from one branch of the model. The Lucas branch — which the same rule reaches at a different growth rate — gives 47 and 76, and neither is a Fibonacci number. The sequence is a consequence of an angle rather than a property of plants.

 What the model settles on, against how fast the meristem grows. A broad golden branch, a transition, and then the two-whorl regime at exactly half a turn. 9 of 27 converged settings land within 4° of the golden angle; 15 land more than 20° away. Where the angle comes from

The bifurcation diagram

Sweep the one parameter of the rule and the settled angle traces a diagram — a broad golden branch, a transition, and a two-whorl regime at exactly half a turn. The famous constant is one branch of it, which is a more useful thing to know than the constant.

 Two runs of the same rule from unrelated starting angles. Both settle at 137.0°, within 0.5° of the golden angle, from seeds 166° apart. Where the angle comes from

Droplets with no biology in them

Douady and Couder dripped magnetised ferrofluid into a dish of silicone oil, and got spiral phyllotaxis with Fibonacci parastichy numbers out of a system containing no cells, no genes and no plant. That is the strongest evidence the pattern is physics — and the clearest warning about what a model can claim.

 What a two-organ cut does at each rise of the 2/3 rung. At every rise the coarse rung is a lattice on, all 36 arrangements of two organs removed, with the ones that never repair counted and split by where they end up. 22 of 324 cuts across the rung reverse the stem's handedness onto the mirror of the divergence they were cut from. 40 fall instead into a cycle whose mean is half a turn, which the lattice they came from has no number for. 4 rises give only the first, 4 give only the second, and at a rise of 0.075 both happen in the same table, which is what says the fate belongs to the cut and not to the rise. Where the angle comes from

Half a turn, four at a time

At four of the nine coarse rises no wrecked cut reverses. What those stems do instead is stop settling: they repeat 171.09°, 269.53°, 189.14°, 90.23° without end, which adds to two whole turns over four organs. The mean is exactly half a turn and a counter finds four files where the lattice had three.

 Two paths down the same tree. Both start at the same first fork. Keeping the larger family every time reaches 137.473°; one different choice reaches 99.549°. Neither angle is in the arithmetic — both are limits of a path. Where the angle comes from

The tree and the attractor

The dynamical model settles on the golden angle over a range of one parameter and on the Lucas angle outside it, and it cannot reach the low-growth end at all. The lattice tree reaches everywhere, has no dynamics in it, and produces the same two angles as limits of two paths. Two routes, one pair of numbers.

 A stem grown at 42 nodes per rung. 167 nodes, each placed where the repulsion from the ones below it was least, with the rise falling from 0.2 to 0.0045. Counted blind in a sliding window the pattern walks 1/2 → 2/3 → 3/5 → 5/8, and the marks are where its answer changed. Where the angle comes from

A pattern with a rate

Every lattice in the essays before this one is a static object indexed by a parameter, and a plant is not. Put the rise on a clock, place each node where the repulsion from the ones below it is least, and the object that comes out has a history — which is the first thing here that could disagree with the ladder.

 The 2-jugate forks converge on 68.7539°. Every fork sits at a rational divergence, with denominator 4(m² + mn + n²) — 10/28, 30/76, 74/196 and so on. The limit is 68.7539°, which is 137.5078 divided by 2, and it is at none of them. The pattern itself

Half the golden angle

The forks of the van Iterson tree converge on 137.5078° and sit at none of them. Divide the whole tree by two and the same thing happens at 68.7539° — which is where teasel is, and where a bijugate sunflower counted 42 and 68 has to be.

 The plane of stems: divergence across, rise up. Each shade is one parastichy pair. The marked points are the lattices where three families are equally short — the forks — and the Fibonacci ones run up the middle towards 137.51°. The claims, measured

The angle is not the object

Every popular account of phyllotaxis is organised around a number. After a round of work spent on stems, forks and frequencies, the number looks like the wrong thing to organise an account around — it is the limit of one path through a branching structure, it is at no fork, and a plant that has it got there by not jumping.

 The disorder of a head against its divergence angle, 300 points. μ₂ is the mean squared departure of a Voronoi cell's side count from six, measured on 300 points inside 86% of the radius. Swept across 1.60° it is a staircase: flat over stretches of a few hundredths of a degree, with sharp steps between them and narrow deep dips wherever a rational falls. At 137.142° — which is 360 × 8/21 — it is 0.078; At 137.646° — which is 360 × 13/34 — it is 0.197; At 138.458° — which is 360 × 5/13 — it is 0.060. The ticks along the top are the fractions p/q, placed from arithmetic rather than from the curve. Packing and tiling

The disorder is a staircase

Sweep the second moment of a head's side-count distribution across the divergence angle and it is not a curve. It is flat in stretches with sharp steps between them and narrow deep dips wherever a rational falls — so 0.253 is not the golden angle's number, it is the number of every angle from 137.47° to 137.54°.

 The dip at 5/13 — 138.4615° — at three head sizes. Walking the divergence angle off an exact rational, at 300, 600, 1200 points. The floor falls as the head grows — 0.059 at 300, 0.029 at 600, 0.014 at 1200, halving for each doubling — and the dip narrows faster: half-widths of 0.0162°, 0.0048°, 0.0010°, a factor of four for each doubling rather than two. Half-width times the square of the head size is 1456, 1726, 1402, which is what makes the width a property of the sample rather than of the angle. Packing and tiling

A dip belongs to the head

At an exact rational the disorder halves when the head doubles, and the dip around it narrows by a factor of four. So which angles look ordered is set by how many organs were counted, and a head of three hundred cannot tell 138.4615° from 138.48° while a head of twelve hundred tells it from 138.4625°.

 One seed, two rates, two ladders. Both stems begin as forty nodes of Lucas lattice at a rise of 0.12. At 65 nodes per rung the divergence stays at 99.5° and the counts walk 1/3 → 3/4 → 4/7 → 7/11. At 131 it leaves for 137.7° and walks 1/3 → 2/3 → 3/5 → 5/8 → 8/13 instead. Where the angle comes from

The rate decides the branch

Forty nodes of Lucas lattice, carried down to the same fine rise twice. Hurried, the pattern holds the Lucas ladder through three more forks at 99.5°. Given room, it abandons it at the first fork it reaches and walks to 8/13 at 137.5°. Same seed, same rise, two ladders — with a threshold between them at about ninety nodes per rung.

 Fibonacci at a rise of 4.8e-3, asked two ways. Choose a divergence at random and a static lattice at this rise gives a consecutive Fibonacci pair 10.8 per cent of the time. Start coarse at a divergence nobody chose, grow the stem down to the same rise, and it is 100 per cent of 16 runs. The geometry is not generous; continuity is. Where the angle comes from

Continuity from a coarse start

At a fine rise, one divergence in seven gives a Fibonacci pair. Grow a stem from a coarse start at a divergence nobody chose, down to that same rise, and sixteen runs out of sixteen end on 8/13. The earlier work's interpretation was that continuity does the work; this is the measurement it never had.

 What the model settles on, against how fast the meristem grows. A broad golden branch, a transition, and then the two-whorl regime at exactly half a turn. 16 of 40 converged settings land within 4° of the golden angle; 17 land more than 20° away. What a plant might be doing

Two-ranked, by two different routes

The rule produces a two-ranked stem at a coarse rise, where 180° is the only thing available, and that has been in the bifurcation diagram since the beginning. It also produces one at a fine rise, at a rise whose own answer is the golden angle, if a single organ is removed. The diagram cannot show the second, and the reason it cannot is how it is drawn.

 The two trees this site draws, at 30° and 32° to a side, against the cost's 37.47° and 37.47°. Two trees of 63 segments each, 5 generations deep and 31 junctions apiece, with every junction's radii taken from r₀³ = r₁³ + r₂³ exactly and every junction's angle taken from a constant. Read as an exponent through cos(θ/2) = 2^(2/p − 1), the drawn angles say 2.5237 and 2.6239, in pictures whose widths are built at exactly 3. The cost that fixed those widths wants 37.47° and 37.47° at this daughter ratio, 74.93° in total, and the misses cost 0.573% and 0.292% of the network — which is why a fixed angle can sit in a figure about a minimisation and never look wrong. Branching and transport

The trees drawn at no angle

Two branching figures in these essays set every junction's radii from the cube law exactly and every junction's angle from a constant nobody derived. Read as exponents the drawn angles say 2.52 and 2.62, in pictures whose widths say exactly three — and at a lopsided fork the drawing puts a daughter thirty-four degrees from where the same cost puts it.

 The disorder of a head against its divergence angle, 900 points. μ₂ is the mean squared departure of a Voronoi cell's side count from six, measured on 900 points inside 86% of the radius. Swept across 1.32° it is a staircase: flat over stretches of a few hundredths of a degree, with sharp steps between them and narrow deep dips wherever a rational falls. At 137.143° — which is 360 × 8/21 — it is 0.025; At 137.882° — which is 360 × 18/47 — it is 0.125; At 138.002° — which is 360 × 23/60 — it is 0.089. The golden angle is marked and sits at 0.253, in the middle of a flat stretch and nowhere near the largest value on the range. The claims, measured

The most irrational is not the most disordered

If rational angles make ordered tissue, the most badly approximable angle should make the most disordered — which would at last give the golden angle a criterion it wins. Swept across the interval it is the most irrational point of, μ₂ peaks at 138.42° and the golden angle sits unremarkably in the middle.

 The fork angle Da Vinci's rule predicts, against the size of the fork, in one tree. A tree has one value of the constant Da Vinci's rule leaves free, so a fork's share of it falls as the square of the fork's size and the angle the rule predicts changes with size. For even forks it is 119.1° at a relative size of 0.1, 101.7° at a relative size of 0.5, 74.9° at a relative size of 1, 44.6° at a relative size of 2, 9.6° at a relative size of 10, where Murray's rule gives 74.93° at every size; for daughter ratio 0.5 it is 118.9° at a relative size of 0.1, 101.3° at a relative size of 0.5, 77.6° at a relative size of 1, 48.9° at a relative size of 2, 10.9° at a relative size of 10, where Murray's rule gives 77.58° at every size. From a total of 100° to 20° at an even fork is a factor of 8.93 in radius. Branching and transport

One constant for every fork

Da Vinci's rule leaves a free constant in the cost that sets a fork's angle, and running it over its range walks the predicted angle from nothing to 120 degrees, through Murray's 74.93. So no single fork can refute the rule. But the constant is one number for a whole tree, and a fork's share of it falls as the square of the fork's size — the constant is a radius axis. A tree spanning a factor of ten in radius must show forks from 29.4 degrees at its biggest to 111.6 at its smallest, a spread wider than one fork's flatness can hide, while Murray's angle is the same at every size.

 Placements that went to a different minimum, per thousand. The rule's own counterfactual, run beside it: where would this node have gone with the noise taken away and everything else left alone? Placement noise displaces the node after the argmin, so the answer is always "here" — 0.2°, 0.4°, 0.8° all give zero. A jostle and a field perturbation are upstream of the choice and change one or two placements in a thousand while the lattice is still intact. Where the angle comes from

Which minimum was chosen

The rule takes an argmin, so there are two completely different things noise can do to it: move the answer, or move the question. One of them can change what is chosen and the other cannot, ever — and the difference is exactly zero against one or two placements in a thousand, at amplitudes where every other measurement says the two are identical.

 The fork angle the transport cost predicts on a tree sized by stress, against the size of the fork. A tree sized so that equal loads on its tips bend every branch to one stress conserves rᵖ with p set by how much shorter each branch is than its parent, λ. On such a tree the cost that fixes a fork's angle carries one constant, entering a fork of size s as s^(2p − 6), so the predicted angle changes with size unless p is three. With the constant set so that a fork of relative size one opens at Murray's 74.9°: At λ = 0.707 (p = 2.000) an even fork opens at 119.1°, 111.6°, 74.9°, 29.4°, 9.6° at relative sizes of 0.1, 0.32, 1, 3.2 and 10; at λ = 0.74 (p = 2.091) an even fork opens at 114.5°, 106.4°, 74.9°, 41.1°, 30.1° at relative sizes of 0.1, 0.32, 1, 3.2 and 10; at λ = 0.794 (p = 2.250) an even fork opens at 106.3°, 97.6°, 74.9°, 53.9°, 46.4° at relative sizes of 0.1, 0.32, 1, 3.2 and 10; at λ = 0.87 (p = 2.498) an even fork opens at 92.7°, 85.5°, 74.9°, 66.0°, 61.5° at relative sizes of 0.1, 0.32, 1, 3.2 and 10; at λ = 0.95 (p = 2.793) an even fork opens at 78.4°, 76.6°, 74.9°, 73.4°, 72.2° at relative sizes of 0.1, 0.32, 1, 3.2 and 10. Murray's rule opens every fork at 74.9°. Branching and transport

Forks on a tree sized by stress

Da Vinci's rule predicts that a tree's forks open wider as they get smaller, because the constant in its angle cost is one number for a tree and enters each fork scaled by its size. A tree sized for equal bending stress conserves an exponent set by how much shorter each branch is than its parent, and on such a tree the same constant enters each fork as its radius to the power 2p − 6. The trend survives at every length ratio short of one and shrinks with it: 105 degrees a decade of radius for a crown filling a plane, 83 at a length ratio of 0.74, 53 for a crown filling a volume, 13 at 0.9. Only a crown shortening about as fast as a volume-filling one fans wider across a tenfold range than one fork's flatness can hide.

 Which rates keep a Lucas seed, on stems of one, two and three organs a whorl, placed by whorls a rung. Every rate each configuration was grown at, drawn at the number of whorls the stem places between one transition and the next: a dark cell keeps the Lucas ladder to the end of the rise, a light cell gives it up. 1 a whorl, 512: kept to 86.6 whorls a rung, lost from 87.6; 1 a whorl, 256: kept to 86.6 whorls a rung, lost from 87.6; 1 a whorl, 1024: kept to 86.6 whorls a rung, lost from 87.6; 2 a whorl, 1024: kept to 87.6 whorls a rung, lost from 88.1; 2 a whorl, 1024, imposed: kept to 87.6 whorls a rung, lost from 88.1; 2 a whorl, 512: kept to 87.6 whorls a rung, lost from 88.5; 2 a whorl, 2048: kept to 87.6 whorls a rung, lost from 88.5; 3 a whorl, 1536: kept to 87.9 whorls a rung, lost from 88.5; 3 a whorl, 1536, imposed: kept to 87.9 whorls a rung, lost from 88.5; 3 a whorl, 768: kept to 86.6 whorls a rung, lost from 88.2. On this axis every edge falls within about two whorls a rung of every other. The pattern itself

A Lucas seed counts whorls

An ordinary stem seeded on the Lucas lattice keeps that ladder when its rise falls fast and gives it up when it falls slowly, with the edge near ninety nodes a rung. A stem that grows two organs a whorl is an ordinary stem folded twice round, so the identity predicts its edge — once it says whether the edge counts placements or whorls. Grown across the edge, stems of one, two and three organs a whorl keep a Lucas seed to 86.6, 87.6 and 87.9 whorls a rung: one number to within a per cent in whorls, and one, two and three times as far out in organs. No grid moves it and imposing exact whorls moves it not at all, even where free trijugate whorls come apart completely as the seed is lost.

 The block is one of the two numbers, and not always the smaller. Every lattice a single organ was removed from, one row each: golden, rise 0.020 carrying 3/5; golden, rise 0.013 carrying 5/8; golden, rise 0.008 carrying 5/8; golden, rise 0.005 carrying 8/13; Lucas, rise 0.020 carrying 4/7; Lucas, rise 0.013 carrying 4/7. The two open ticks on each line are that lattice's own parastichy numbers; the filled dots are the blocks the stems that never recovered settled into, one per offset that failed. The claim this table was built to test is that the block is the smaller of the two, which held at the two rises it was first measured at. It does not hold here: 3/5 gives 5, 5/8 gives 5 and 8, 4/7 gives 4 and 7. What survives is weaker and still worth something — every filled dot but 1 sits on one of that row's own ticks, so the orbit carries a count of the lattice it was cut from, and which of the two it carries is decided by the offset rather than by the pattern. Stems and cones

Two accounts of one number

A stem that never recovers from an ablation settles into a repeating block whose length was the smaller of its two spiral counts, at both arrangements it had been measured at. Two different explanations predicted exactly that and could not be told apart. Swept across four rises on the ordinary branch the premise itself fails: at one arrangement the block is the larger number, at another both appear, and the rule that seemed to be there was two measurements.

 One rule, one rise, two branches that stay where they were put. The top 70 organs of two stems grown by the same placement rule at the same rise of 0.013, differing only in the stretch of ideal lattice each was started from. The left one was seeded at the golden angle and settles at 136.781° with the pair 5/8; the right one was seeded on the Lucas lattice and settles at 99.785° with 4/7. Neither drifts towards the other: 0.73° and 0.28° from where each was seeded, over four hundred organs. That is what makes an intervention on the right-hand stem a measurement about a different lattice rather than about a different rule — and 4 and 7 are not Fibonacci numbers, which is the property the experiment needs. Stems and cones

A stem on the other branch

Every stem an organ had been cut from carried Fibonacci counts, which is why two rival explanations of the block a wrecked stem settles into had never disagreed. A stem seeded on the Lucas lattice carries four and seven at the same rise, under the same rule. Cut, it settles on seven — the larger number, and not a Fibonacci one.

 Every angle whose counts from 34 to 144 resist approximation within a per cent of the golden angle's. Over the counts a head shows from 34 to 144, an angle scores like the golden angle when those counts add up, each the sum of the two before, from a first pair near the golden ratio. 46 angles between 20° and 180° come within one per cent of its score of 0.44718, each drawn as a stem at its angle. The five nearest are 137.51° with counts 34, 55, 89, 144 at 100.000 per cent; 99.50° with counts 47, 76, 123 at 99.989 per cent; 106.45° with counts 44, 71, 115 at 99.931 per cent; 151.14° with counts 50, 81, 131 at 99.919 per cent; 132.18° with counts 49, 79, 128 at 99.907 per cent. The golden angle is the highest, and the Lucas angle at 99.50° is a ten-thousandth of the score behind it. The claims, measured

What a head can mean by most irrational

Hurwitz's bound, the one famous claim about this subject that survives, is a limit over every denominator, and a head shows only the counts between its innermost spirals and its rim. Over those counts an angle resists approximation like the golden angle exactly when the counts it shows add up, each the sum of the two before, from a pair near the golden ratio — and every such pair has an angle of its own. The golden angle still scores highest over every window measured, by a ten-thousandth: over counts from 34 to 144 the Lucas angle is 99.989 per cent of it and forty-six angles are within one per cent. What separates the golden angle from them is below the counts they share, at the centre of the head.

 Fork angles on crowns sized by the larger of flow and stress, handing over at six different generations. Thirteen-generation crowns at a length ratio of 0.707, every branch sized by the larger of flow and stress, twigs at the radius the transport cost prefers, with the two radii equal at generations 3 to 8. Handing over at 3, the forks from the trunk open at 38, 51, 67, 75°; handing over at 4, the forks from the trunk open at 27, 38, 51, 67, 75°; handing over at 5, the forks from the trunk open at 19, 27, 37, 51, 67, 75°; handing over at 6, the forks from the trunk open at 13, 18, 26, 36, 50, 66, 75°; handing over at 7, the forks from the trunk open at 8, 12, 17, 24, 35, 49, 66, 75°; handing over at 8, the forks from the trunk open at 4, 6, 10, 15, 22, 32, 46, 65, 75°. Every fork from the handover outward opens at Murray's 74.9°. No fork on any of the six opens wider than that, and none turns back: the angle rises through the stress-sized generations and stops. Branching and transport

A trend that stops at Murray's angle

A crown sized by whichever of flow and bending stress asks for the thicker branch is sized by stress at its trunk end and by flow at its twigs, and the twigs fix the constant that was free in the fork-angle prediction: a twig at the radius the transport cost prefers spends exactly half its upkeep on pumping. On such a crown the fork angle does not change sign at the handover. It rises through every stress-sized generation, meets Murray's 74.93° at the handover and stays there, and no fork anywhere opens wider. The trend turns back only when the twigs are thinner than the cost wants — past a pumping share of (λ^(−2/3) − 1)/(1 − λ^(4/3)), 0.70 at the planar crown and closing on one half as branches stop shortening.

 On a rung the response is a run of offsets; near a transition it has a hole. One row per rise, from 0.02 at the top to 0.005 at the bottom, and one column per offset: the organ one place back at the left, 14 places back at the right. A cell is filled where removing that organ moves the next organ by more than 2.5°, and empty where it does not. On a rung the filled cells are a run from one to the larger parastichy number — 8, 12, 13 at the pairs shown on the left. Between a rise of 0.02 and 0.0065 the run ends at 5 and one more cell is filled at 7, with the offsets between them quiet to under a degree. That isolated column is one place inside the larger number of the pair the stem is climbing towards. Where the angle comes from

The response with a hole in it

Removing an organ is felt out to the larger parastichy number and no further — that is the intervention's headline, and it holds in the middle of a rung. Swept towards a transition the run of felt offsets stops early and one lone offset past it comes alive, with three quiet organs in between. The lone offset is one place inside the count the stem is about to have.

 The block a wrecked stem settles into is the count it was cut from. A stem is cut in the middle of its front and followed for 300 organs. It does not come back to its lattice; what it does instead is repeat a fixed sequence of divergences exactly, to the resolution of the azimuth grid. Each row shows that sequence twice over, one bar per organ, drawn to the same scale. At the 5/8 rung the sequence is five angles long and advances -36.1° per block; At the 8/13 rung the sequence is eight angles long and advances 22.5° per block. Every block is the smaller number of the pair that was cut, so the second attractor carries the first one's count — and every precession is one part in twice the block, which makes each of these a two-jugate arrangement with 10 and 16 rows. Where the angle comes from

The block is the count it was cut from

A stem that never recovers from an ablation settles into a repeating block of eight angles precessing by 22.7°. Eight was also the smaller parastichy number of the lattice that was cut, which left two possibilities and no way to choose between them. Cut a stem one rung coarser and the block is five.

 The wrecked stem is the lattice it was cut from, wound the other way. The divergence of each organ placed after two were removed from a settled stem at a rise of 0.032, over the 220 organs following the cut. The upper line is the divergence the undisturbed stem holds, 139.6875°; the lower is 220.3125°, which is 360° minus it and therefore the same lattice with the opposite handedness. The sequence is thrown by the cut, wanders for a few dozen organs, and settles on the second line to four decimal places — 220.3125° against 220.3125° — where it stays. A counter shown the positions afterwards returns 3 and 5, the pair the stem was cut from. Nothing about the pattern has been lost; its chirality has been reversed, which at this rung a single removal cannot do. Where the angle comes from

The stem that changed hands

A stem that never recovers from an ablation is supposed to end up somewhere worse than it started — a repeating block of angles, a pattern with the wrong counts in it. At the coarsest arrangement it ends up somewhere that is not worse at all: at 220.3125°, which is 360° minus the divergence it was cut from. The lattice is intact and its handedness is reversed.

 The settled divergence down the golden branch. Every rise from 0.07 down to 0.00482, plotted against the divergence the rule settles on, with each rung drawn in its own stroke and the branch's limit angle marked. The curve does not slide: it turns three times in four rungs, climbing across one and falling across the next, so a value it takes on one rung it takes again on another. That is what makes a matched pair possible — two rises, different counted pairs, one angle — and it is the whole reason the design exists on this branch. The widest excursions from the limit angle, coarse rung first, are 3.195°, 3.352°, 0.961°, 0.422°. Stems and cones

The angle the ladder returns to

Down the golden branch the settled divergence climbs across one rung and falls across the next, turning three times in four rungs. That is why the same angle is reached at two different rises — and why the Lucas branch, which turns once, almost never offers the same thing.

 Six boundaries on three basins, five kinds between them, and three that are edges. Each basin's stretch of starting angle, with both its boundaries located to ± 0.125° by sweeping ten degrees at a quarter of a degree. The pale bar behind each is the interval the forty-angle table could bracket it in, three to five degrees at a time. The widest basin, at rise 0.03 and exponent 2, is a fringe at 124.375° and a puncture at 179.625°. The same-rise basin, at rise 0.03 and exponent 3, is a sliver at 124.875° and a fade at 167.625°. The narrow basin, at rise 0.02 and exponent 3, is a wall at 144.875° and a fade at 161.625°. Only 3 of the 6 are edges in the sense of a side: the widest basin's upper boundary is a hole 0.75° wide centred on 180°, with the same destination beyond it, so that basin runs out of basin at the reflection point rather than reaching an edge. The interval the forty-angle table bracketed each basin in is drawn behind it, from the sweep at 1200 organs a run. What a plant might be doing

A basin with no upper edge

The widest basin in the settling table had a width bracketed between 47.5 degrees and about 57, and closing a bracket means sampling near an edge rather than everywhere. Three basins cut at a quarter of a degree located all six of their boundaries, and the widest turned out to run out of basin at 180 degrees rather than reach an edge on that side at all.

 Every divergence a stem settles on, by falloff exponent. One row per exponent, one mark per distinct settled divergence anywhere in that column, read to a tenth of a degree. This is the sharpest instrument in the sweep and the cheapest: it is already in every run, it does not average, and it is read on the same azimuth grid at every exponent. The steeper falloffs reach 42.3°, 47.9°, 148.1° — values that appear nowhere in the two shallower columns — while the traffic the other way is empty. So the exponent changes the map of where a stem can end up while leaving alone the share of starting angles that end up anywhere, which is a result a measurement of "did it settle" was never going to find. Where the angle comes from

Destinations only a steep rule reaches

A settled stem's divergence is the sharpest instrument in the sweep and the cheapest: it is already in every run, it does not average, and it is read on one grid at every exponent. Exponents 4 and 5 reach 42.3°, 47.9° and 148.1°, and the two shallower ones reach none of them.

 Every destination in the settling table, counted. One row per destination, placed across at the divergence the stems settled on and labelled with the parastichy pair a counter returns at the top of the run. 117 settled runs land on 15 destinations when two values within half a degree are called one. five of them sit on the golden or the Lucas sequence and ten do not, and the ones that do not are reached at every falloff exponent including the shallowest. Nothing here refuses to count: the settling test does not admit arrangements this collection would decline to call lattices. Where the angle comes from

A counter on the settling table

The settling table has reported an angle for every run that reaches a lattice, and nobody has ever counted one. A hundred and seventeen settled runs, regrown and counted: every one of them has a parastichy pair, and sixty-five of them land on sequences the ladder does not carry.

 What the 3 steep-only destinations count as. Each block is a divergence that only the two steeper falloff exponents settle a stem on, with the pair a counter returns and the sequence that pair belongs to. All three count. None is a rung of either ladder. Two of them are the only members of their sequence anywhere in the table; the third, at 148.1 degrees, is the coarsest rung of a sequence that supplies another destination at every exponent — so it is not a new kind of arrangement but a coarser one of a kind the table already reached. Where the angle comes from

What a steep rule counts as

Exponents four and five settle stems on 42.3°, 47.9° and 148.1°, and the two shallower ones reach none of them. All three count: 8/9, 8/15 and 2/5. None is a rung of either ladder, and one of them is the coarsest rung of a sequence the table already had.

 The 10 sequences the settling table's destinations belong to. Every pair the counter returns is two consecutive terms of a sequence in which each term is the sum of the two before it, which is the ladder's own rule. two of these are the sequences plants are observed to follow. The rest are not, and they are not rare: the 1, 4, 5, 9, 14 sequence and the 2, 5, 7, 12, 19 sequence each supply several destinations, at every falloff exponent the table is grown at. The right-hand column is the divergences that land on each. Where the angle comes from

Ten sequences, two of them the ladder's

Every parastichy pair the settling table produces is two consecutive terms of a sequence in which each term is the sum of the two before it. Ten such sequences account for all fifteen destinations — nine, once one of the ten turns out to be a reading rather than a ladder — and the two the collection is built on are neither the largest nor the smallest.

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