Topological charge — the series
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An interior that is nearly neutral
Give every cell a charge of six minus its number of sides and the total over a tessellated head is fixed by its own boundary, exactly, with nothing left over for the interior. On a golden head that freedom is spent on 264 exceptions among 1,631 cells which cancel to six.
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The defects lie on rings
The cells in a seed head that are not hexagons are not scattered through it. Every one of 264 sits within a cell of a radius computed from the divergence angle alone, the radii are a factor of φ apart, and between two of them lie 422 consecutive cells without a single exception.
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Every five is bound to a seven
A five-sided cell beside a seven-sided one is one object in a crystal and two exceptions in a tiling, and the phyllotaxis literature borrows the crystallographic word without measuring the binding. Measured against a seeded permutation null on a 2,400-organ head, every five in the interior shares a wall with a seven, and the loneliest one in the head is 0.927 cell spacings from the nearest.
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The rings are not the transitions
A seed head has two ladders on it — the radii where the counted parastichy pair changes, and the radii where the exceptional cells sit — and the obvious guess is that they are the same ladder. They are not: the second sits a factor of the square root of phi outside the first at every rung of two different divergence ladders, which is exactly halfway between two consecutive transitions.
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A dislocation that closes
A defect ring on a seed head looks exactly like a low-angle grain boundary: a closed chain of bound five–seven pairs one cell thick. The defining test of a dislocation is a loop that fails to close. On an elastic crystal it fails by one lattice vector at every size. On a golden head's rings the tightest loop round a five–seven pair closes every time, 272 pairs out of 272, and one set of lattice labels holds every patch across the outer rings without a single broken edge. No lattice rotates at the ring, where a real boundary of that density would need thirty-two degrees. What a ring actually is sits in one number: its walls are within two over its own count of being replaced by the other diagonal of their cell, on one curve for all four rings.