Fibonacci — where it appears
Named by 52 essays across 7 fields — each of them below, with the objects they name alongside it.
The angle is an output
137.5° is not a constant of nature. It is where a rule settles — a rule that places each new element as far as it can from the ones already there, contains no reference to the golden ratio, and reaches the same answer from starting angles a hundred and sixty degrees apart.
Fibonacci is a branch, not a law
Fibonacci counts come from one branch of the model. The Lucas branch — which the same rule reaches at a different growth rate — gives 47 and 76, and neither is a Fibonacci number. The sequence is a consequence of an angle rather than a property of plants.
Counting the spirals
Almost every claim about phyllotaxis is a claim about how many spirals run through a pattern, and the count is almost never done. It can be done from the points alone, by a count that is never told what angle built them — and then a count of 34 is evidence rather than a restatement.
Counting up the stem
The same counting machinery, pointed at a stem instead of a seed head, returns one answer three times where the head returned three answers. That contrast is a measurement rather than a preference, and it is the one the whole cylindrical argument rests on.
Fitting the exponent
Assuming the exponent is three and reporting the error says how far the data is from that assumption. Fitting the exponent and reporting what it comes out as says what the network is doing — and an estimator has to be shown returning something other than three, or it is not a fit.
The bifurcation diagram
Sweep the one parameter of the rule and the settled angle traces a diagram — a broad golden branch, a transition, and a two-whorl regime at exactly half a turn. The famous constant is one branch of it, which is a more useful thing to know than the constant.
The claim that survives
Of the three famous assertions about this subject, one is out by a factor of two, one is true of a branch rather than of plants, and one is right — in a sharper form than the version usually told, and about arithmetic rather than about packing.
The counts change with radius
The same head gives 13 and 21 near the centre, 21 and 34 further out, 34 and 55 beyond that, and 55 and 89 at the rim. The transitions are at computable radii, and a photograph captioned with one pair is a statement about one annulus rather than about a flower.
Two numbers out of the points
A seed head's divergence angle can be recovered from its spiral counts only to within an interval, because a range of angles gives the same counts. On a stem the counts come with lengths attached, two measurements pin two unknowns, and the lattice comes back to the last digit it was built with.
What "whorled" was hiding
The earlier work's census put 35% of divergences in a bucket labelled whorled and moved on. Opened, every pair in it is k and 2k — the coarsest rung of the ladder, repeated k times — and reading the census up to jugacy takes the Fibonacci share from 14.7% to 50.1% without describing a single extra plant.
The nautilus question
A golden spiral grows by 6.854 per turn. Measured nautilus sections give about 3.2. That is a factor of 2.14 — not a rounding error, not an artefact of where the centre is assumed to be, and not close.
Recovering the angle from the counts
Build a head at a stated divergence angle, forget the angle, and get it back from the spiral counts alone. Four angles, worst error twelve thousandths of a degree — and the only thing that crossed between the two halves was a list of coordinates.
The Fibonacci ladder
Lower the rise on a cylinder and the parastichy pair climbs — 1 and 2, then 2 and 3, then 3 and 5 — each rung the sum of the two before it. Nothing in the arithmetic mentions Fibonacci, the transitions sit at computable rises, and consecutive ones stand in the ratio 1/φ².
Where the model stops
Below a growth parameter of about 0.18 this implementation does not converge — the settled angle wanders over a hundred degrees however long the run. That is the range where the literature says the interesting behaviour lives, and it is worth a figure rather than a quietly chosen axis.
The tree and the attractor
The dynamical model settles on the golden angle over a range of one parameter and on the Lucas angle outside it, and it cannot reach the low-growth end at all. The lattice tree reaches everywhere, has no dynamics in it, and produces the same two angles as limits of two paths. Two routes, one pair of numbers.
The forks are exact
Where a stem's pattern has to choose between two futures, three spiral families are equally short and the lattice is exactly equilateral. A numerical solver found those points; the numbers it returned turned out to be rational, and chasing that gave a closed form — including the fact that every fork sits at a rational divergence, and the golden angle at none of them.
Half the golden angle
The forks of the van Iterson tree converge on 137.5078° and sit at none of them. Divide the whole tree by two and the same thing happens at 68.7539° — which is where teasel is, and where a bijugate sunflower counted 42 and 68 has to be.
The angle is not the object
Every popular account of phyllotaxis is organised around a number. After a round of work spent on stems, forks and frequencies, the number looks like the wrong thing to organise an account around — it is the limit of one path through a branching structure, it is at no fork, and a plant that has it got there by not jumping.
Transitions a factor of φ² apart
The ladder's rungs are a factor of 1/φ² apart in rise. A disc's rise falls as one over radius squared and a cone's as one over distance, so the same rungs land a factor of φ apart on a seed head and a factor of φ² apart on a cone — measured, on both, by a counter that has never heard of either.
What a count is worth
A reported parastichy pair pins the divergence angle to a band 221°/mn wide — so 2 and 3 says almost nothing and 34 and 55 fixes it to a tenth of a degree. Each step up the Fibonacci sequence is worth a factor of φ², and recording the radius a pair was counted at adds only ten per cent.
Every family but two is a sum
A seed head has six spiral families and everybody reports two. That looks like a convention hiding information and it is the opposite — every family but the two smallest is the sum of two others, so a third count is a prediction rather than a measurement, and a check that catches a wrong pair.
The rate decides the branch
Forty nodes of Lucas lattice, carried down to the same fine rise twice. Hurried, the pattern holds the Lucas ladder through three more forks at 99.5°. Given room, it abandons it at the first fork it reaches and walks to 8/13 at 137.5°. Same seed, same rise, two ladders — with a threshold between them at about ninety nodes per rung.
How many plants would it take
Fourteen specimens separate the geometry's Fibonacci share from a coin weighted to a half. Four separate it from what a grown history gives. One fir cone measured at three rings settles whether its transitions are spaced as a cone's or an ogive's. The sample sizes are small, and that is the uncomfortable part.
Continuity from a coarse start
At a fine rise, one divergence in seven gives a Fibonacci pair. Grow a stem from a coarse start at a divergence nobody chose, down to that same rise, and sixteen runs out of sixteen end on 8/13. The earlier work's interpretation was that continuity does the work; this is the measurement it never had.
The angles name the branch
Seed the same rule at the Lucas angle and the readout returns 4 and 7, then 7 and 11 — the pairs the position counter finds, and not Fibonacci numbers. So a list of divergence angles carries not only how many spirals there are but which family of ladders the plant is on.
The survey this site cannot do
Four rounds of asking for a dataset, and it is still not here. What the work here can do instead is specify it — the fields, the sampling, the sizes, and which of this collection's claims each one would settle. Two of the four fields asked for turn out to be worth less than the asking implied, and one was never asked for at all.
Noise is not a slow rate
A stem seeded on the Lucas branch keeps it below ninety nodes per rung and abandons it above — which invites the objection that a real apex's fluctuations would knock it off regardless. Measured across a hundred and sixty runs of two independent kinds of noise, one escapes, at the amplitude where the pattern is already coming apart.
How many organs a pair needs
A count taken over too few organs does not fail. It returns the rung below, which is a perfectly good pair, and nothing anywhere says so. The window that avoids it is not a constant but the counter's own arithmetic, and 384 settled runs sit exactly where that arithmetic puts them.
A count set by a delay
An L-system describes a plant and forbids nothing, because none of its parameters is anything a plant has. One branching grammar is the exception: a mature apex makes a new bud every season, and a bud waits d seasons before it branches. Its counts grow at the root of x^(d+1) = x^d + 1, a delay of one season gives Fibonacci's numbers and nothing else does, and the fourth count already separates a one-season wait from every longer one. So a Fibonacci count in a branching plant is a measurement of how long its buds wait. It is also a fragile one: if one bud in ten waits two seasons instead, eleven counts in a row come out Fibonacci's three times in a thousand.
A count that loses its growing points
The branching grammar behind the Fibonacci claim has no deaths in it, and a stem that loses shoots is the common case. Giving every growing point a chance q of dying each season leaves the counts a linear recurrence and does exactly one thing to it: the growth rate becomes the deathless root multiplied by 1 − q, at every delay and every death chance, to the last bit a double holds. So each waiting time has a death chance above which its lineage shrinks — a half with no wait, 0.3820 at one season, 0.2451 at four — and a longer wait tolerates less. What does not survive is the count itself: a plant losing one growing point in ten a season shows eight Fibonacci counts in a row one time in ten thousand, against one time in eight for a bud that occasionally waits an extra season.
What a scar is worth
Counting the scars a dead shoot leaves does not put a branching count back on the sequence it would have had. A scar records a growing point and a dead growing point takes every branch it would have made, so living points plus scars reach 39.2 per cent of the deathless count after twenty seasons at one death in twenty, and 1.9 per cent at one in five — falling without limit rather than closing. What the scars restore is the other number. Scars per living point settle at q/(x − 1) exactly, so a rate with a scar share beside it recovers the death chance and then the waiting time, where a rate alone is reached by a one-season wait losing a tenth, a two-season wait losing 0.64 per cent and no wait at all losing 27.2 per cent.
A head displaced before it is counted
The round trip from a head's spiral counts back to its divergence angle was tested on heads whose every organ sat exactly where the rule put it. Displaced by a normal error of up to two and a half spacings, heads of 900 organs keep counting a pair from their own sequence and return intervals holding the true angle to a spacing and a half; heads of 300 organs move to the neighbouring pair by half a spacing and then refuse, nine in ten of them by two spacings. Every moved count brings in the family whose chord was third shortest. Of 898 heads recovered, 19 intervals miss the true angle and 17 of those by about a tenth of a degree — displacement makes the reading coarser and then silent, not confidently wrong.
A count that can be wrong by one
A reported parastichy pair pins the divergence angle to a band 221°/mn wide only if both counts are right. Allowing either to be off by one adds the bands of every neighbouring pair whose counts share no factor, and those bands sit where their own lattices live: for 2/3 they swallow the whole range, and for 34/55 they are two bands as narrow as the true one at 109° and 113°, twenty-five degrees away. So a high count that may be wrong is not a blurred reading but a short list of sharp candidates, costing log₂ 3 bits. And on the Fibonacci pairs, two in every six — 21/34 and 34/55 among them — cannot be miscounted silently by one count at all, because every such miscount shares a factor.
A twist is a divergence
Recovering a head's divergence angle from its spiral counts survived independent displacements of whole spacings, moving to a neighbouring pair and then refusing rather than misleading. Displacements with a direction are harder on it in only one case. A head pressed to an aspect ratio of 2.25, spread at the rim by eighty per cent or sheared with a slope of 1.6 is still counted as its own pair or its neighbour, and recovered inside its interval. A head twisted — each organ turned about the centre in proportion to its radius — is not: past a turn of the rim of about a radian and a half the counts leave their sequence and the recovered angle misses, by up to sixty-one degrees, because a twist changes the angle between one organ and the next. The round trip is not fooled. It is reporting the angle the twisted head has.
Seven rises and two seeds
One organ removed from a stem is felt out to the larger of its two spiral counts. Every test of that has confounded the count with the rise, because on one branch the two move together. Grow a second branch beside the first at the same rise and they come apart — and doing it at seven rises turns a matched pair into a design whose last column changes hands four times.
What a head can mean by most irrational
Hurwitz's bound, the one famous claim about this subject that survives, is a limit over every denominator, and a head shows only the counts between its innermost spirals and its rim. Over those counts an angle resists approximation like the golden angle exactly when the counts it shows add up, each the sum of the two before, from a pair near the golden ratio — and every such pair has an angle of its own. The golden angle still scores highest over every window measured, by a ten-thousandth: over counts from 34 to 144 the Lucas angle is 99.989 per cent of it and forty-six angles are within one per cent. What separates the golden angle from them is below the counts they share, at the centre of the head.
The count sees the twist first
A twisted head recovers a changed divergence, and the check proposed for it was two annuli: the twist's extra angle falls with radius, so an inner and an outer annulus should disagree. Read on golden heads of 900, 2,400 and 9,000 organs, they never do in time. Their intervals separate at eight radians on 900 organs and never on the larger heads, always after the ordinary reading has been misled — from six radians on 900 organs and from two on 2,400 and 9,000. What catches the twist first, at every size and on every seed, is the count: at half a radian to three quarters some band stops returning two consecutive Fibonacci numbers — 34 and 89, 89 and 233 — which no untwisted golden head, clean or displaced, ever does.
A count that drifts by two
A reported pair of 34 and 55 that may be wrong by one allows three sharp bands; allowed to drift by two it allows thirteen, and by three, twenty-nine — and the information lost is exactly the logarithm of that count, because every band is as narrow as the true one. The nearest wrong band stays twenty-four degrees away until a drift of three brings one to eleven. What does not survive is the protection: 21/34 and 34/55 could not be miscounted silently by one, but every Fibonacci pair can be by two, so a counter who drifts by two as readily as by one reports 34/55 silently wrong 9.5 per cent of the time rather than 0.13.
The flag reads the angle, not the twist
A twisted seed head is caught first by its counts: at half a radian some band stops returning two consecutive Fibonacci numbers, which an untwisted golden head never does. Twist the head in proportion to the square of the radius instead and the organs land exactly where a head grown at the golden angle plus a/(N − 1) puts them — to a billionth of a spacing — so the two read the same pairs in every band, and the flag fires on both. The flag detects a divergence that is not golden, and it has a resolution: 0.015° on 900 organs, 0.01° on 2,400, 0.003° on 9,000. Every twist shape from a half to four is flagged exactly when its change to the divergence in the outer annulus passes that resolution. So a flagged head is not golden, and nothing in its counts says whether it was twisted or grown that way.
The hole on the other branch
Near a transition, the run of offsets a stem notices stops being a run: there is quiet past the front and then one isolated offset, felt as hard as anything inside it. Where that offset sits was pinned down on Fibonacci lattices, where the numbers to check it against are 5, 8 and 13. On the Lucas branch they are 4, 7 and 11 — and the rule holds there too.
Two ways to die, three things to count
Giving a branching plant's waiting buds a death chance of their own leaves its counts a linear recurrence, but breaks the collapse onto the survival: the rate becomes the apex survival times the root of y^(d+1) = y^d + r^d, where r is the bud survival over the apex survival. The one-chance reading then names the wrong waiting time on 171 of 477 plants with waits of two to four seasons, shorter when the buds are the fragile ones and longer when the apices are. The two chances are separable from a rate and a scar share, exactly — but the two counts' loci cross at eight to sixteen degrees, so a one per cent error lets the chances wander by a factor of two. A third count is owed, and it is the scars sorted by kind.
A bad year does not average out
Let every growing point on a plant share one season's death chance, bad one season in ten and good otherwise, with the average held at 0.1. Averaged over every sequence of seasons the counts are exactly the independent model's. But no plant is an average over sequences: with bad years at 0.5 a plant settles on a rate of 1.3001 against the expected 1.3190, the median plant holds 54 per cent of the expected count by season forty, and the scar share never settles, because it is set by how many seasons ago the last bad year was — 1.35 the season after one, 0.17 twelve seasons on. So the reading of a plant's waiting time from a rate and a scar share gets worse the longer it runs: right for 57 per cent of plants over ten seasons, 22 per cent over eighty.
Two counts that slip together
A counter who closes the circle a few degrees late counts a sliver of the head twice, in both families at once, so the two counts of a reported pair drift together rather than apart. Coupled that way the count is safer than it was: fourteen Fibonacci pairs in twenty-three admit no silent equal shift of one, against seven that admit no silent single miscount, and 34/55 announces every closing error short of 9.82°. The check is what breaks. Two annuli closed at the same wrong mark pass 17.6 per cent of wrong readings of 34/55 and 76.8 per cent of 13/21's, because a linear relation survives multiplication — and what catches them instead is a protractor good to twelve degrees.
The census wants a low count
Four specimens separate the geometry's Fibonacci share of 14.7 per cent from the ninety per cent a grown history gives — if every count is right. Counted with a closing error spread over 7.2°, the same census needs six specimens counted at 13/21, fifty-four at 34/55 and 449 at 55/89, because the geometry's own pairs are all small enough that no closing error under 11° moves them, while a grown plant counted high loses its Fibonacci reading first. Counted at 55/89 with a spread of 9.83° the census reads plants as less Fibonacci than random angles. The count that pins the divergence best is the one a census should avoid.
Scars with dates on them
A bad season shared by every growing point wrecks the reading of a branching plant by two totals: over eighty seasons the rate and the scar share name a two-season wait for 22 per cent of plants. Date the scars — by position along a shoot, by growth ring — and each season's death chance is read off its own scars, so the bad years stop being noise and become a known input. Running the branching recurrence through the plant's own seasons names the wait for 94 per cent of plants over twenty seasons whether bad years kill a tenth of the points or seven tenths, and for every plant over eighty. What a plant cannot read from its own scars is the climate: the averaged chance comes only as fast as seasons do.
Two marks chosen by one eye
A counter traces each family of spirals from a starting organ of its own, so a reported pair carries two closing errors, correlated because one eye chose both. Letting them differ costs 34/55 its ten-degree margin — 33/56 and 35/54 share no factor, and marks that err 5.3° in opposite directions reach them — while 21/34 keeps its margin whatever the marks do. And it decides the second annulus. At a spread of 7.2° the relation passes right readings 2.8 times as readily as silent ones when the marks are independent, 1.25 times at a correlation of 0.9, and stops telling them apart at 0.98; where it does work it keeps one reading in forty-six.
Counting it again
A reading whose two counts share a factor says the count went wrong, and the specimen is still there to be counted again. Counted afresh, the reading kept is exactly one reading conditioned on not announcing itself — the second chance a silent error gets is matched by the second chance a right reading gets — so a recount changes which specimens a census keeps, not what a kept reading says. At 34/55 with closing errors spread over 7.2° it takes the census from fifteen kept specimens to ten and from about thirty counts to twenty-one, and against scoring every reading it turns 449 counts at 55/89 into 52. It never makes a high count as cheap as counting 13/21 once.
The rung decides the sign
Twenty-four lattices where both walls of the slot are really there. Thirteen give a strongly positive interaction, at 85° to 135°; eleven give a negative or null one, at −25° to −0.5°. Nothing lies between. Every rung's lattices fall on the same side as each other.
A counter on the settling table
The settling table has reported an angle for every run that reaches a lattice, and nobody has ever counted one. A hundred and seventeen settled runs, regrown and counted: every one of them has a parastichy pair, and sixty-five of them land on sequences the ladder does not carry.
What a steep rule counts as
Exponents four and five settle stems on 42.3°, 47.9° and 148.1°, and the two shallower ones reach none of them. All three count: 8/9, 8/15 and 2/5. None is a rung of either ladder, and one of them is the coarsest rung of a sequence the table already had.
Ten sequences, two of them the ladder's
Every parastichy pair the settling table produces is two consecutive terms of a sequence in which each term is the sum of the two before it. Ten such sequences account for all fifteen destinations — nine, once one of the ten turns out to be a reading rather than a ladder — and the two the collection is built on are neither the largest nor the smallest.
A list that can only shrink
The destinations only a steep falloff reaches grew from three to five when the sampling doubled, and everyone read it as a list filling in. Doubling again takes two off it, which is the only direction a list defined by an absence can ever move.
Named alongside it
The objects these essays reach for when they reach for this one.
Divergence angleHonest limitsBranchLadderRiseParastichy pairClaim testingLucas numbersMeasurementIdentifiabilityMeasurement errorRound trip