Concept

Fibonacci — where it appears

25 essays name this object, across 6 fields. What follows is each of them, and the objects they name alongside it.
golden 137.51°55 and 89FibonacciLucas 99.50°47 and 76Lucas151.14°50 and 81neither77.96°37 and 60neithercounted from the pointsone sequence per branch

Fibonacci is a branch, not a law

Fibonacci counts come from one branch of the model. The Lucas branch — which the same rule reaches at a different growth rate — gives 47 and 76, and neither is a Fibonacci number. The sequence is a consequence of an angle rather than a property of plants.

wrong · fibbranch
05e+51e+62e+60100200300angle around the boundary (°)repulsion from what is theregrowth 0.40 · 14 elements in playthe minimum is where the next one goes

The angle is an output

137.5° is not a constant of nature. It is where a rule settles — a rule that places each new element as far as it can from the ones already there, contains no reference to the golden ratio, and reaches the same answer from starting angles a hundred and sixty degrees apart.

emergence · attractor
21 and 34 spiralscounted, not assumed

Counting the spirals

Almost every claim about phyllotaxis is a claim about how many spirals run through a pattern, and the count is almost never done. It can be done from the points alone, by machinery that is never told what angle built them — and then a count of 34 is evidence rather than a restatement.

lattices · counting
stem, lower third2 and 3stem, middle third2 and 3stem, upper third2 and 3disc, r = 0.18–0.3221 and 34disc, r = 0.45–0.6234 and 55disc, r = 0.82–0.9955 and 89stem at rise 0.050, disc of 1600 points, both at 137.51°counted from coordinates onlyone answer against three

Counting up the stem

The same counting machinery, pointed at a stem instead of a seed head, returns one answer three times where the head returned three answers. That contrast is a measurement rather than a preference, and it is the one the whole cylindrical argument rests on.

cylinder · stem count
02468234exponent khow badly r₀ᵏ = Σ rᵢᵏ fails at that kk = 3127 junctionsfitted k = 3.000

Fitting the exponent

Assuming the exponent is three and reporting the error tells you how wrong the data is. Fitting the exponent and reporting what it comes out as tells you what the network is doing — and the machinery has to be shown returning something other than three, or it is not a fit.

branching · exponent
1001251501750.50011.50growth parameter Gangle the model settles on (°)goldentwo-whorlfilled: converged · hollow: still wanderingone rule, one knob

The bifurcation diagram

Sweep the one parameter of the rule and the settled angle traces a diagram — a broad golden branch, a transition, and a two-whorl regime at exactly half a turn. The famous constant is one branch of it, which is a more useful thing to know than the constant.

emergence · bifurcation
00.2000.400100120140160divergence angle (°)resistance to rational approximation (higher is more irrational)1/√5 — the bound137.508° — 0.438938 angles on a 0.08° grid, plus the golden angle exactlythis claim is sharp

The claim that survives

Of the three famous assertions about this subject, one is out by a factor of two, one is true of a branch rather than of plants, and one is right — in a sharper form than the version usually told, and about arithmetic rather than about packing.

wrong · hurwitz
0255075406080fraction of the head's radiusparastichy numbers found in a band there1000 primordia at the golden angle4 different pairs

The counts change with radius

The same head gives 13 and 21 near the centre, 21 and 34 further out, 34 and 55 beyond that, and 55 and 89 at the rim. The transitions are at computable radii, and a photograph captioned with one pair is a statement about one annulus rather than about a flower.

lattices · transitions
golden — 6.85× per turnnautilus — 3.2× per turnsame construction, same start2.14× apart in growth

The nautilus question

A golden spiral grows by 6.854 per turn. Measured nautilus sections give about 3.2. That is a factor of 2.14 — not a rounding error, not an artefact of where the centre is assumed to be, and not close.

shells · nautilus
137.51°, rise 0.098.5e-14°counted 2/3137.51°, rise 0.032.0e-13°counted 3/5137.51°, rise 0.0122.8e-14°counted 5/899.50°, rise 0.083.0e-13°counted 1/3151.14°, rise 0.071.1e-13°counted 2/399.50°, rise 0.021.1e-13°counted 4/7error in the recovered divergence anglecounts and hop lengths onlyworst 3.0e-13°

Two numbers out of the points

A seed head's divergence angle can be recovered from its spiral counts only to within an interval, because a range of angles gives the same counts. On a stem the counts come with lengths attached, two measurements pin two unknowns, and the lattice comes back to the last digit it was built with.

cylinder · cylinder recovery
5/107.9%5 × 1/23/67.9%3 × 1/22/46.1%2 × 1/24/85.5%4 × 1/27/145.2%7 × 1/26/122.8%6 × 1/28/160.1%8 × 1/2share of divergences at this riserise 0.008 · 3600 divergences35.4% share a factor

What "whorled" was hiding

The expansion phase's census put 35% of divergences in a bucket labelled whorled and moved on. Opened, every pair in it is k and 2k — the coarsest rung of the ladder, repeated k times — and reading the census up to jugacy takes the Fibonacci share from 14.7% to 50.1% without describing a single extra plant.

wrong · jugate census
the first of the four — 300 of its 899 pointsused to buildcountsrecovered137.508°55 · 89137.520°99.502°47 · 7699.500°151.100°31 · 81151.105°77.960°37 · 6077.960°worst error 0.012°counts in, angle outthe recovery never sees the angle

Recovering the angle from the counts

Build a head at a stated divergence angle, forget the angle, and get it back from the spiral counts alone. Four angles, worst error twelve thousandths of a degree — and the only thing that crossed between the two halves was a list of coordinates.

lattices · recovery
0.2500.5000.75011.25-2.50-2-1.50-1-0.500log₁₀ of the rise between nodes (falling to the right is the plant growing)log₁₀ of the larger parastichy number2/33/55/88/1313/21500 rises, shortest vectors recomputed at eachratio 0.3820 against 1/φ² = 0.3820

The Fibonacci ladder

Lower the rise on a cylinder and the parastichy pair climbs — 1 and 2, then 2 and 3, then 3 and 5 — each rung the sum of the two before it. Nothing in the arithmetic mentions Fibonacci, the transitions sit at computable rises, and consecutive ones stand in the ratio 1/φ².

cylinder · ladder
0501000.2500.5000.7501growth parameter Gspread of the last 30 steps (°) — 0 means settledfilled dark: convergedthe usable range is stated, not implied

Where the model stops

Below a growth parameter of about 0.18 this implementation does not converge — the settled angle wanders over a hundred degrees however long the run. That is the range where the literature says the interesting behaviour lives, and it is worth a figure rather than a quietly chosen axis.

emergence · modellimit
-2-1.50-1-0.500100120140160180divergence angle (°)log₁₀ of the rise between nodes1,2,32,3,522 × 150 lattices, each solved196 runs drawn

The forks are exact

Where a stem's pattern has to choose between two futures, three spiral families are equally short and the lattice is exactly equilateral. A numerical solver found those points; the numbers it returned turned out to be rational, and chasing that gave a closed form — including the fact that every fork sits at a rational divergence, and the golden angle at none of them.

cylinder · forks
100120140-3-2-1log₁₀ of the rise at the forkdivergence angle at the fork (°)137.508° — Fibonacci99.502° — Lucas13 forks, each solved for three equal families137.4730° and 99.5495°

The tree and the attractor

The dynamical model settles on the golden angle over a range of one parameter and on the Lucas angle outside it, and it cannot reach the low-growth end at all. The lattice tree reaches everywhere, has no dynamics in it, and produces the same two angles as limits of two paths. Two routes, one pair of numbers.

emergence · branch tree
646668701234567fork number down the treedivergence at the fork (°)137.5078/2 = 68.7539°2-jugate · 7 forkslast fork 68.7365°

Half the golden angle

The forks of the van Iterson tree converge on 137.5078° and sit at none of them. Divide the whole tree by two and the same thing happens at 68.7539° — which is where teasel is, and where a bijugate sunflower counted 42 and 68 has to be.

lattices · jugate limit
-2-1.50-1-0.500100120140160180divergence angle (°)log₁₀ of the rise between nodes1,2,32,3,522 × 150 lattices, each solved196 runs drawn

The angle is not the object

Every popular account of phyllotaxis is organised around a number. After a phase spent on stems, forks and frequencies, the number looks like the wrong thing to organise an account around — it is the limit of one path through a branching structure, it is at no fork, and a plant that has it got there by not jumping.

wrong · the tree not the angle
-3-2-112distance from the apex, log₁₀rise in local circumferences, log₁₀flare 0.35 · step 15 transitions, ratio 2.619

Transitions a factor of φ² apart

The ladder's rungs are a factor of 1/φ² apart in rise. A disc's rise falls as one over radius squared and a cone's as one over distance, so the same rungs land a factor of φ apart on a seed head and a factor of φ² apart on a cone — measured, on both, by a counter that has never heard of either.

cylinder · cone transitions
-10111.5022.503product of the two counts, log₁₀angles left open, log₁₀ °7 pairs · edges found by bisectionwidth × mn = 221°

What a count is worth

A reported parastichy pair pins the divergence angle to a band 221°/mn wide — so 2 and 3 says almost nothing and 34 and 55 fixes it to a tenth of a degree. Every rung of the ladder is worth a factor of φ², and the counting radius this collection has asked published counts for since its foundation adds ten per cent.

wrong · count value
00.2500.5000.75015101520specimenschance of detecting it14 specimens90%exact binomial · one-sided at α = 0.0514 specimens, cut at 5

How many plants would it take

Fourteen specimens separate the geometry's Fibonacci share from a coin weighted to a half. Four separate it from what a grown history gives. One fir cone measured at three rings settles whether its ladder is spaced as a cone's or an ogive's. The sample sizes are small, and that is the uncomfortable part.

wrong · sample size
10011012013014011.502falling rise, as −log₁₀divergence the stem is producing (°)137.51°, Fibonacci99.50°, Lucasseeded at 99.50°, rise 0.127/11 against 8/13

The rate decides the branch

Forty nodes of Lucas lattice, carried down to the same fine rise twice. Hurried, the pattern holds the Lucas ladder through three more forks at 99.5°. Given room, it abandons it at the first fork it reaches and walks to 8/13 at 137.5°. Same seed, same rise, two ladders — with a threshold between them at about ninety nodes per rung.

emergence · branch choice
grown from a coarse start100.0%divergence chosen at random10.8%share ending on a consecutive Fibonacci pair16 grown runs, starting divergences from 67° to 299°every one of them ended on 8/1367 nodes per rung · rise 0.4 → 4.8e-3100% against 10.8%

Continuity from a coarse start

At a fine rise, one divergence in seven gives a Fibonacci pair. Grow a stem from a coarse start at a divergence nobody chose, down to that same rise, and sixteen runs out of sixteen end on 8/13. The expansion phase's interpretation was that continuity does the work; this is the measurement it never had.

emergence · continuity
plants show consecutive Fibonacci pairs far more…4and more often even than a coin weighted to a half14a conifer cone's rings are spaced as a cone rather…1multijugate patterns are a real minority rather than…34against 14.7%, if the truth is 90%needs: the pair, at a stated rungagainst 14.7%, if the truth is 50%needs: the pair, at a stated rungagainst φ² = 2.62, if the truth is φ^(2/1.88) = 1.67needs: three ring positions, to ±3%against 2%, if the truth is 15%needs: the pair; the whorl's symmetryspecimens neededexact binomial · α = 0.05 · power 0.91 to 34 specimens

The survey this site cannot do

Four phases of asking for a dataset, and it is still not here. What this phase can do instead is specify it — the fields, the sampling, the sizes, and which of this collection's claims each one would settle. Two of the four fields asked for turn out to be worth less than the asking implied, and one was never asked for at all.

wrong · survey spec
placement noisedegrees off the minimum00.250.50.7511.251.52field noisefraction of the barrier00.00250.0050.00750.010.01250.0150.02kept its branchchanged branchanother pairno latticeseeded 40 nodes of Lucas lattice · 10 runs per amplitude1 escape in 160 runs

Noise is not a slow rate

A stem seeded on the Lucas branch keeps it below ninety nodes per rung and abandons it above — which invites the objection that a real apex's fluctuations would knock it off regardless. Measured across a hundred and sixty runs of two independent kinds of noise, one escapes, at the amplitude where the pattern is already coming apart.

emergence · noise amplitude

Named alongside it

The objects these essays reach for when they reach for this one.

BranchRiseDivergence angleLadderCylinderφ, the golden ratioRepulsionAttractorCensusParastichySurveyTransitions

All concepts