Phyllotaxis
the spirals are counted, not admired
Concept

Vogel's model — where it appears

The construction that places the nth floret at the golden angle times n and a radius proportional to the square root of n. It is a description of a finished head rather than an account of how one is made, and it fits by assuming the answer.

Named by 6 essays across 3 fields — each of them below, with the objects they name alongside it.

A head of 200 primordia at a divergence of 137.51°. Nothing is placed by hand: the nth point sits at n·137.51° and radius √n. The closest any two points come is 1.60 of the mean spacing.

A head is a set of points

The nth primordium at n times an angle, and a radius of root n. Two lines of arithmetic produce a sunflower head, which is either remarkable or suspicious depending on how carefully the claim is stated — and stating it carefully is most of the work.

lattices · Lattice Where the disc's counts change, predicted from a cylinder. The dashed lines are the transition radii the cylinder's ladder gives through h = c²/4πr², with nothing fitted. The dots are what the blind counter returns from the disc: 15 of 16 bands agree, and the ones that do not sit on a transition.

A disc is a cylinder

Vogel's seed head makes the rise fall as one over radius squared, so a disc is not one lattice but a family of them. Feed that into the cylinder's ladder and it predicts where a sunflower's spiral counts change — with nothing fitted, and against a counter that never sees either model.

cylinder · Rise and radius 135 fives and 129 sevens among 1631 interior cells. The side counts of every cell strictly inside a golden, 137.508° head of 2400 organs, cut at 86% of the radius. The fives and the sevens are counted apart rather than summed into a spread, because they are opposite charges and the sum hides them. Summed over the interior, 264 cells that are not hexagons carry a charge of +6. Over the whole patch the charge is 294, which is exactly 6 + 2·144 — a number fixed by the 144 cells on the patch's own boundary and carrying nothing whatever about the interior. The defects are not scarce; they are balanced.

An interior that is nearly neutral

Give every cell a charge of six minus its number of sides and the total over a tessellated head is fixed by its own boundary, exactly, with nothing left over for the interior. On a golden head that freedom is spent on 264 exceptions among 1,631 cells which cancel to six.

tissue · Topological charge Every five is a cell away from a seven, and the loneliest is 0.927 spacings out. How far a five-sided cell is from the nearest seven-sided one, in cell spacings, on a golden, 137.508° head of 2400 organs. The measured bar runs from the closest five to the loneliest — 0.833 to 0.927, with a median of 0.919. It stops a single cell out, so there is no unpaired tail at all rather than a small one. The nulls are seeded permutations over 200 draws: relabelling which defects are fives puts the average five 1.254 ± 0.064 spacings away, and scattering the whole multiset over the interior cells puts it 1.786 ± 0.092. 100.0% of the fives share a wall with a seven against 68.2% for the strong null, z = 7.3.

Every five is bound to a seven

A five-sided cell beside a seven-sided one is one object in a crystal and two exceptions in a tiling, and the phyllotaxis literature borrows the crystallographic word without measuring the binding. Measured against a seeded permutation null on a 2,400-organ head, every five in the interior shares a wall with a seven, and the loneliest one in the head is 0.927 cell spacings from the nearest.

tissue · Topological charge The cell area a 137.508° golden head's packing is divided by, at 10 sizes. Counting every bounded cell the mean area runs from 3.607 to 38.828, the largest at 150 organs, because the cells just inside the edge of the head reach out to circumcentres far beyond it. Leaving out every cell whose polygon crosses the head's own radius, it stays between 3.1425 and 3.1634 at every size drawn, which is π, the area the square-root rule gives each organ.

Packing, measured against the interior

An earlier reading of these heads reported that no packing criterion singles out the golden angle and that three criteria give three winners. Every one of those readings was divided by a mean cell area that, on a head of 150 organs, was 38.8 where the interior's is π. Divided by the interior's own, the criteria about distance put the golden angle first of 72 angles and the criteria about cells go to rational ones.

tissue · Packing How much a twist of the rim changes the divergence between neighbouring organs, across a 900-organ head, for six shapes of twist. A twist of 0.25 radians at the rim, shaped as a·(r/R)^p, changes the angle between one organ and the next by a·(p/2)·ρ^(p−2)/(N − 1) at a distance ρ from the centre: the six lines are exponents 0.5, 1, 1.5, 2, 3, 4. At two the change is the same everywhere — 0.0159° — which is a different divergence angle; below two it is largest at the centre, above two at the rim. The shaded ring is the outer annulus, and the band between dotted lines is the flag's resolution on an untwisted 900-organ head: grown 0.0125° off the golden angle it counts consecutive Fibonacci numbers, 0.015° off it does not. At this twist the outer annulus passes it for exponents 2, 3, 4.

The flag reads the angle, not the twist

A twisted seed head is caught first by its counts: at half a radian some band stops returning two consecutive Fibonacci numbers, which an untwisted golden head never does. Twist the head in proportion to the square of the radius instead and the organs land exactly where a head grown at the golden angle plus a/(N − 1) puts them — to a billionth of a spacing — so the two read the same pairs in every band, and the flag fires on both. The flag detects a divergence that is not golden, and it has a resolution: 0.015° on 900 organs, 0.01° on 2,400, 0.003° on 9,000. Every twist shape from a half to four is flagged exactly when its change to the divergence in the outer annulus passes that resolution. So a flagged head is not golden, and nothing in its counts says whether it was twisted or grown that way.

lattices · Recovery

Named alongside it

The objects these essays reach for when they reach for this one.

Divergence angleHonest limitsRim effectCylinderDelaunayParastichyParastichy pairRational angleTopological chargeArtefactBound pairCell area

All concepts