Phyllotaxis
the spirals are counted, not admired
Concept

Meristem growth — where it appears

The expansion of the tip itself while organs are being placed on it, which changes the geometry each new organ is placed into. Treating a growing organ as a term in the rule rather than as a fixed point is a separate hypothesis with its own consequences.

Named by 4 essays across 2 fields — each of them below, with the objects they name alongside it.

The rule, 26 steps in, at a growth of 0.40. The next primordium goes where the repulsion is least — the marked minimum at 216°. Nothing in the rule refers to any particular angle.

The angle is an output

137.5° is not a constant of nature. It is where a rule settles — a rule that places each new element as far as it can from the ones already there, contains no reference to the golden ratio, and reaches the same answer from starting angles a hundred and sixty degrees apart.

emergence · Attractor Three disturbances, three places to get in. The rule reads its neighbours, builds a profile of the energy at every azimuth, takes the least of it, and records a position. field noise enters at the profile; jostle noise enters at the neighbours; placement noise enters at the record. Two of the three are upstream of the choice and can change which minimum is taken; the third is downstream and never can.

The noise that arrives through the neighbours

The two kinds of noise this site had were idealisations that bracket the rule's choice. The realistic disturbance is neither: a primordium is placed exactly, and then the organ grows, so by the time the next one forms its neighbours have moved. That is a third kind, and it is invisible in every measurement a plant offers.

mechanism · Noise entry Three disturbances, three places to get in. The rule reads its neighbours, builds a profile of the energy at every azimuth, takes the least of it, and records a position. jostle noise enters at the neighbours; placement noise enters at the record. Two of the three are upstream of the choice and can change which minimum is taken; the third is downstream and never can.

A growing organ is part of the rule

Every model in the earlier essays places primordia on a surface and then treats the surface as furniture. But the surface grows between one placement and the next, and that growth reaches the rule through the only channel it has — where the neighbours are. What looks like a boundary condition turns out to be a term in the model.

mechanism · Mechanism claims A window that fits inside a rung. Stems that climb the ladder at four rates, read over a window at the fine end. The condition is a ratio: the window has to be shorter than a rung. 250 internodes at 130 per rung is 1.92 rungs and agrees on 0 of 3; 400 internodes at 130 per rung is 3.08 rungs and agrees on 0 of 3; 250 internodes at 260 per rung is 0.96 rungs and agrees on 3 of 3; 400 internodes at 260 per rung is 1.54 rungs and agrees on 1 of 3; 250 internodes at 520 per rung is 0.48 rungs and agrees on 2 of 3; 400 internodes at 520 per rung is 0.77 rungs and agrees on 3 of 3; 250 internodes at 1040 per rung is 0.24 rungs and agrees on 3 of 3; 400 internodes at 1040 per rung is 0.38 rungs and agrees on 3 of 3. Read over the whole stem instead, every rate returns nothing — 0 of 3, 0 of 3, 0 of 3, 0 of 3 — because the quantity the comb is periodic in changes as the pattern climbs.

A window inside a rung

A stem that climbs the ladder has no comb in it at any rate, because the quantity the comb is periodic in changes as it goes. Read a window instead and it comes back, on one condition: the window has to be shorter than a rung — which makes the shoot's rate the thing that decides whether a plant can be asked.

emergence · Rising

Named alongside it

The objects these essays reach for when they reach for this one.

Divergence angleBasinElongationEpitheliumHonest limitsIdentifiabilityMeasurementMechanismMeristemNoiseThe placement rulePrimordium

All concepts