Phyllotaxis
the spirals are counted, not admired
Concept

Basin — where it appears

The set of starting conditions from which a dynamic reaches a given attractor. Which lattice a stem ends on is a question about which basin its seed sat in, so a claim that one pattern is selected is a claim about basins and not about optima.

Named by 28 essays across 3 fields — each of them below, with the objects they name alongside it.

Three disturbances, three places to get in. The rule reads its neighbours, builds a profile of the energy at every azimuth, takes the least of it, and records a position. field noise enters at the profile; jostle noise enters at the neighbours; placement noise enters at the record. Two of the three are upstream of the choice and can change which minimum is taken; the third is downstream and never can.

The noise that arrives through the neighbours

The two kinds of noise this site had were idealisations that bracket the rule's choice. The realistic disturbance is neither: a primordium is placed exactly, and then the organ grows, so by the time the next one forms its neighbours have moved. That is a third kind, and it is invisible in every measurement a plant offers.

mechanism · Noise entry Three disturbances, three places to get in. The rule reads its neighbours, builds a profile of the energy at every azimuth, takes the least of it, and records a position. jostle noise enters at the neighbours; placement noise enters at the record. Two of the three are upstream of the choice and can change which minimum is taken; the third is downstream and never can.

A growing organ is part of the rule

Every model in the earlier essays places primordia on a surface and then treats the surface as furniture. But the surface grows between one placement and the next, and that growth reaches the rule through the only channel it has — where the neighbours are. What looks like a boundary condition turns out to be a term in the model.

mechanism · Mechanism claims What the sequence sees that the scatter cannot. Each point is an ensemble at one amplitude, placed at the scatter it produces. A stem at three quarters of a degree of scatter has a lag-one correlation near zero if its noise arrived after the primordium was placed, and near 0.7 if it arrived before — and no measurement of the scatter can tell those apart. The separation closes above about a degree, because what the other two kinds preserve is the correlation of a lattice.

What one angle says about the next

A tenth of a degree of placement noise moves a stem's divergence scatter from 0.50° to 0.62°, which nobody would report. It takes the correlation between consecutive angles from 0.54 to below zero. The other two kinds of noise, at scatters where no measurement can separate them, leave it at 0.6.

cylinder · Sequence Placements that went to a different minimum, per thousand. The rule's own counterfactual, run beside it: where would this node have gone with the noise taken away and everything else left alone? Placement noise displaces the node after the argmin, so the answer is always "here" — 0.2°, 0.4°, 0.8° all give zero. A jostle and a field perturbation are upstream of the choice and change one or two placements in a thousand while the lattice is still intact.

Which minimum was chosen

The rule takes an argmin, so there are two completely different things noise can do to it: move the answer, or move the question. One of them can change what is chosen and the other cannot, ever — and the difference is exactly zero against one or two placements in a thousand, at amplitudes where every other measurement says the two are identical.

emergence · Noise entry Where each kind's lattice gives way. The largest amplitude at which every run still has a lattice, and the scatter it produces there. The amplitudes are incomparable — field 0.015 (fraction of the barrier), jostle 1 (degrees of azimuth), placement 0.8 (degrees of azimuth) — and the scatters agree to 19%. The boundary belongs to the pattern rather than to the disturbance: a lattice fails at about a degree and a half of scatter, and which of three mechanisms produced it does not move where.

The boundary belongs to the pattern

Three kinds of noise, in three incommensurable units, destroy a lattice at the same place — about a degree and a half of divergence scatter. The earlier work measured that of two kinds and called it a scale rather than a constant. With a third it looks less like a coincidence and more like a property of what a lattice is.

emergence · Scatter tolerance The wrecked stem is the lattice it was cut from, wound the other way. The divergence of each organ placed after two were removed from a settled stem at a rise of 0.032, over the 220 organs following the cut. The upper line is the divergence the undisturbed stem holds, 139.6875°; the lower is 220.3125°, which is 360° minus it and therefore the same lattice with the opposite handedness. The sequence is thrown by the cut, wanders for a few dozen organs, and settles on the second line to four decimal places — 220.3125° against 220.3125° — where it stays. A counter shown the positions afterwards returns 3 and 5, the pair the stem was cut from. Nothing about the pattern has been lost; its chirality has been reversed, which at this rung a single removal cannot do.

The stem that changed hands

A stem that never recovers from an ablation is supposed to end up somewhere worse than it started — a repeating block of angles, a pattern with the wrong counts in it. At the coarsest arrangement it ends up somewhere that is not worse at all: at 220.3125°, which is 360° minus the divergence it was cut from. The lattice is intact and its handedness is reversed.

emergence · Attractor How many organs a stem needs before it is on a lattice. One row per rise, one mark per starting angle, placed at the organ from which every later divergence stays within a degree and a half of the run's own final value. Where a stem settles at all it does so between 0 and 290 organs in, against the 400 every ablation run here grows before it cuts anything. Not one row needs the length it is given. What changes down the table is the count on the right: how many of the nine starting angles reach a lattice at all, which falls from 7 at the coarse rises to 1 at the finest.

How long a stem takes to settle

Every result here is grown on a stem that has settled onto a lattice, and settling has always been tested for and never timed. Timed, it takes between nothing and two hundred and ninety organs — against the four hundred every ablation run grows before it cuts anything, and the nine hundred the noise runs carry.

cylinder · Settling The same table, grown 2.7 times as long. Every rise and every starting angle, grown to 1200 organs and then to 3200. The two middle columns are how many starting angles reached a lattice at each length, and they are the same column: of the 72 pairs of runs, 72 are identical organ for organ and 0 settle at the longer length after failing at the shorter one. Tripling the budget buys nothing anywhere. What the fine rises are short of is not run length: the share of starting angles that reach a lattice at all falls from 7 of 9 to 1, so the arrangements a stem could fall into have mostly stopped existing.

A wall and not a budget

Below a rise of about 0.005 this collection's stems stop settling onto a lattice, and the limit has been written up four times without anybody asking which kind of limit it is. Grown three times as long, the table is identical row for row: not one stem that failed to settle succeeds. The floor is a wall.

cylinder · Settling The mirror belongs to the lattice, not to the dose. How close the closest arrangement came to the mirror of the divergence it was cut from, against the share of the front that was removed. The marked point at 40 per cent is the coarse 3/5 rung with two organs taken, which reaches the mirror exactly. Every other point is a finer rung: five sizes of cut at 5/8 running from 13 to 63 per cent, and three organs at 8/13. Taking a larger share of a larger front than the coarse rung needs gets nowhere near, so the quantity that decides it is not the fraction of the neighbourhood removed.

The share was not the thing

Two organs out of a front of five reverses a stem's handedness; three out of eight does not, and neither does five out of eight, which is a larger share of a larger neighbourhood. The hypothesis under test was that the dose decides the destination. It decides whether a stem falls off its lattice and nothing about where it lands.

emergence · Dose The short list, and what is on it. Every distinct place a wrecked stem settles, for cuts of one organ through five at one rung, with what each one turned out to be. Three of the six are the lattice the stem was cut from with one lag left rigid and a whole number of turns inserted over its period — the description the single-organ work established. The rest have no rigid lag at any period up to twenty-four, and a counter shown their positions returns a pair the original lattice does not carry. So the list is short and it is not homogeneous: a large enough cut can put a stem onto a different lattice rather than onto a slipped version of its own.

A wreck has a short list

Cuts of one organ through five, over two hundred and forty-six stems that never came back, land on six settled divergences between them. Removing five organs instead of one wrecks nearly everything and reaches nowhere the single cut had not already found — and half the list turns out to be the old lattice slipped by a turn, while the other half is not the old lattice at all.

emergence · Dose Six boundaries on three basins, five kinds between them, and three that are edges. Each basin's stretch of starting angle, with both its boundaries located to ± 0.125° by sweeping ten degrees at a quarter of a degree. The pale bar behind each is the interval the forty-angle table could bracket it in, three to five degrees at a time. The widest basin, at rise 0.03 and exponent 2, is a fringe at 124.375° and a puncture at 179.625°. The same-rise basin, at rise 0.03 and exponent 3, is a sliver at 124.875° and a fade at 167.625°. The narrow basin, at rise 0.02 and exponent 3, is a wall at 144.875° and a fade at 161.625°. Only 3 of the 6 are edges in the sense of a side: the widest basin's upper boundary is a hole 0.75° wide centred on 180°, with the same destination beyond it, so that basin runs out of basin at the reflection point rather than reaching an edge. The interval the forty-angle table bracketed each basin in is drawn behind it, from the sweep at 1200 organs a run.

A basin with no upper edge

The widest basin in the settling table had a width bracketed between 47.5 degrees and about 57, and closing a bracket means sampling near an edge rather than everywhere. Three basins cut at a quarter of a degree located all six of their boundaries, and the widest turned out to run out of basin at 180 degrees rather than reach an edge on that side at all.

mechanism · Attractor Two instruments at six boundaries: the clock cannot tell a wall from a fade and the tail spread separates them by a factor of 327. The clock is the ratio of the slowest settling in the last degree inside a basin to the middle of the rest of its window. It rises towards every boundary and diverges at none: the one clean wall rises by 1.313× and the two clean fades by 1.00× and 1.57×, so the wall sits between them and no threshold on a clock separates the two kinds. The tail spread is how far a run's own last two hundred organs wander, and every run has one whether it settles or not. Past the wall it is 0.101°, as steady as the basin just left; past the two fades it is 33.042° and 39.712°. Pooled over all 574 runs everything that settles spreads by 0–0.442° and everything that does not by 30.828–39.712°. The two instruments are drawn side by side on one row per boundary, from the sweep at 1200 organs a run.

A wall or a fade

A basin's border is either a change of destination or a stretch where the angles stop settling at all, and nothing here could tell the two apart. Two instruments were pointed at the question: the settling clock, which looked obviously right and fails, and the tail spread, which was already being computed on every run and had never been read.

mechanism · Attractor 1.75° of starting angle between two basins that reaches neither, and a 3° void beside it. 20.25 degrees of starting angle swept unbroken at 0.25°, from inside the basin at 101.5° to inside the widest basin at 139.3°, one cell per sampled angle. The last angle reaching 101.5° is 114.5° and the first reaching 139.3° is 116.25°; the 1.75° between them holds 6 sampled angles and 0 of them settle anywhere at all. The widest run of angles reaching nothing is 3° wide, at 118–120.75°, and a detached island of 139.3° sits between the two. Starting angle is left over. The stretch that reaches neither basin is bracketed above the strip, from the sweep at 1200 organs a run.

The angles left over

A stem started anywhere on the circle was assumed to end up in one basin or another, so that the settled destinations divided the starting angles between them. Twenty and a quarter degrees swept without a hole at a quarter of a degree find 1.75 degrees between two basins that reaches neither of them and nothing else, and a three-degree void beside it.

mechanism · Attractor How many starting angles reach a lattice, by rise and by falloff. Every cell is 9 stems grown from 9 starting angles spanning 40° to 180°, at one rise and one exponent in the placement rule's falloff, for 1200 organs each. The number is how many of them settle onto a lattice. Reading down a column, the basin closes as the rise falls, which is the wall. Reading across a row, nothing much happens: the exponent-three column is the published settling table digit for digit, and the other three are the same column inside their own sampling error. The account that a steeper falloff should wall somewhere else does not survive the table it predicted.

A steeper rule walls nowhere else

The account of the wall at the fine end was that the basin narrows because the neighbourhood deepens, which predicts a steeper falloff walling somewhere else. Grown at four exponents, the four columns settle 30, 31, 29 and 27 of 72 — a spread of 0.056 against an error of 0.058.

emergence · Settling 41.75° in the middle of the widest basin, sampled only at the table's own 3.12–4.38°. The widest basin from its located lower boundary at 124.375° to the reflection point at 180°, with the two ten-degree windows swept at 0.25° shaded and the stretch between them left open. Nothing has looked inside that stretch more finely than the forty-angle table's own 3.12–4.38° spacing. The narrowest feature this sweep found anywhere is the 0.75° wedge inside same-rise, and a feature that size falls between the table's angles 79% of the time — 75% under an even 3° sampling. So an unmeasured sliver or puncture could sit anywhere in the middle of this basin and nothing here would have seen it. The two swept windows are shaded and the stretch between them is left open, from the sweep at 1200 organs a run.

What a quarter degree cannot see

Six boundaries were located to an eighth of a degree, three basins were named and one width was quoted, and every one of those readings has the same floor under it. The sweep's grid is one step of the grid the stems are placed on, so nothing here bounds a basin narrower than half a degree — and the widest basin's own middle was never swept at all.

mechanism · Attractor How many organs settling takes, by falloff exponent. One mark per stem that settles, placed at the organ from which every later divergence stays within a degree and a half of the run's own final value. At the exponent every other measurement here uses, the slowest is 290 organs — the number four rounds of this collection have carried as though it belonged to the rule. It belongs to the exponent: at 4 the slowest is 808 and at 5 it is 674. A table of shares cannot see this, because a stem that settles at organ 800 and one that settles at organ 8 are the same entry in it. And it is still not a shortage of time: of 72 pairs of runs grown to 1200 organs and then to 3200, 0 settle at the longer length after failing at the shorter.

The clock a share cannot see

Settling takes nothing to 290 organs, and four rounds of this collection have carried that as a property of the rule. At a steeper falloff the slowest is 808 — and not one of the 72 pairs of runs settles at 3,200 organs after failing at 1,200, so the fine end is still a wall.

emergence · Settling Every divergence a stem settles on, by falloff exponent. One row per exponent, one mark per distinct settled divergence anywhere in that column, read to a tenth of a degree. This is the sharpest instrument in the sweep and the cheapest: it is already in every run, it does not average, and it is read on the same azimuth grid at every exponent. The steeper falloffs reach 42.3°, 47.9°, 148.1° — values that appear nowhere in the two shallower columns — while the traffic the other way is empty. So the exponent changes the map of where a stem can end up while leaving alone the share of starting angles that end up anywhere, which is a result a measurement of "did it settle" was never going to find.

Destinations only a steep rule reaches

A settled stem's divergence is the sharpest instrument in the sweep and the cheapest: it is already in every run, it does not average, and it is read on one grid at every exponent. Exponents 4 and 5 reach 42.3°, 47.9° and 148.1°, and the two shallower ones reach none of them.

emergence · Settling Every destination in the settling table, counted. One row per destination, placed across at the divergence the stems settled on and labelled with the parastichy pair a counter returns at the top of the run. 117 settled runs land on 15 destinations when two values within half a degree are called one. five of them sit on the golden or the Lucas sequence and ten do not, and the ones that do not are reached at every falloff exponent including the shallowest. Nothing here refuses to count: the settling test does not admit arrangements this collection would decline to call lattices.

A counter on the settling table

The settling table has reported an angle for every run that reaches a lattice, and nobody has ever counted one. A hundred and seventeen settled runs, regrown and counted: every one of them has a parastichy pair, and sixty-five of them land on sequences the ladder does not carry.

emergence · Settling What the 3 steep-only destinations count as. Each block is a divergence that only the two steeper falloff exponents settle a stem on, with the pair a counter returns and the sequence that pair belongs to. All three count. None is a rung of either ladder. Two of them are the only members of their sequence anywhere in the table; the third, at 148.1 degrees, is the coarsest rung of a sequence that supplies another destination at every exponent — so it is not a new kind of arrangement but a coarser one of a kind the table already reached.

What a steep rule counts as

Exponents four and five settle stems on 42.3°, 47.9° and 148.1°, and the two shallower ones reach none of them. All three count: 8/9, 8/15 and 2/5. None is a rung of either ladder, and one of them is the coarsest rung of a sequence the table already had.

emergence · Settling The 10 sequences the settling table's destinations belong to. Every pair the counter returns is two consecutive terms of a sequence in which each term is the sum of the two before it, which is the ladder's own rule. two of these are the sequences plants are observed to follow. The rest are not, and they are not rare: the 1, 4, 5, 9, 14 sequence and the 2, 5, 7, 12, 19 sequence each supply several destinations, at every falloff exponent the table is grown at. The right-hand column is the divergences that land on each.

Ten sequences, two of them the ladder's

Every parastichy pair the settling table produces is two consecutive terms of a sequence in which each term is the sum of the two before it. Ten such sequences account for all fifteen destinations — nine, once one of the ten turns out to be a reading rather than a ladder — and the two the collection is built on are neither the largest nor the smallest.

emergence · Off-ladder The destinations a steep rule reaches, read two ways. Above: the list read to a tenth of a degree, which is what the settling table prints — 13 entries. Below: the same list read at half a degree, which is two steps of the grid the azimuths are placed on — 3. The 10 entries the finer reading adds are destinations a shallow exponent also reaches, a tenth or two of a degree away, and a tenth of a degree is a fifth of one step of that grid. The pale marks are every destination in the table, for scale.

A list that was a rounding

Three destinations only a steep falloff reaches, read at two grid steps. Thirteen, read at a tenth of a degree. And four arrangements, read by what the counter returns rather than by the angle — a different four, with one the angle reading hides.

emergence · Off-ladder Where a stem started at each of twenty angles ends up, at a falloff exponent of 3. One column per starting angle and one row per rise, coarse at the top. A filled cell is a run that reached a lattice, its tone the destination it reached; a pale cell is a run that never settles. Runs of one tone across neighbouring columns are basins, and the widest of them spans 7 consecutive angles. The angles are 6.25 to 10 degrees apart, so a basin narrower than that cannot be seen here and a single filled cell says nothing about how wide its basin is.

A basin has a width

A destination reached from one starting angle is a presence. A destination reached from seven consecutive starting angles spanning forty-five degrees is a basin with an extent, and nine angles could not have measured one — they were too far apart to have two of them land in the same place.

emergence · Settling The settling share at every rise, sampled two ways. How often a stem reaches a lattice, against the rise, at each of the four falloff exponents. The horizontal rule is the half share the wall is read at. At nine starting angles the four exponents cross it at rises spanning a factor of 1.96 and order themselves 2, 5, 3, 4; at twenty they span a factor of 1.18 and order themselves 5, 4, 2, 3. The ordering reverses and the spread collapses, so the conclusion that the exponents do not separate is confirmed and the numbers that had suggested otherwise were the coarser sampling's own error.

A wall that stopped moving

Four falloff exponents were reported not to move the rise below which stems stop reaching a lattice. Their measured walls spanned a factor of two and ordered themselves 2, 5, 3, 4. At twenty starting angles they span a fifth of one and order themselves 5, 4, 2, 3 — so the conclusion was right and its arithmetic was noise.

emergence · Exponent How often each group of starting angles reaches a lattice. The share of runs that settle, for the nine angles the table was grown from, for eight angles placed halfway between them, and for three angles below the nine's lowest. The nine settle far more often than either. So it is the refinement rather than the extension that drags the share down, and the settling share this collection reports is biased upwards by the choice of angles rather than by their range.

Round numbers are not a sample

The nine starting angles the settling table was grown from reach a lattice four times in ten. Eight angles placed exactly halfway between them reach one a quarter of the time. The difference is not noise and it is not the range — several of the nine sit next door to somewhere a stem could settle.

emergence · Settling

Twenty angles instead of nine

Every claim in this collection about where a stem ends up rests on nine starting angles a cell, and the file that uses them says so — it computes a binomial error of 0.17 and declines to read a spread against it. Eleven more angles halve that error and change what several of the numbers were.

emergence · Settling

A wrecked run goes somewhere

Where a wrecked stem finishes was called unstable. Half of them finish within a degree of a destination measured from intact stems started at arbitrary angles — two tables that share no run, no design and no question.

cylinder · Both walls

Forty angles, and a limit

Nine starting angles turned out to be a biased sample of the circle, and doubling to twenty said by how much. Doubling again says the estimate is converging — to a smaller correction than one doubling extrapolated to.

emergence · Settling

A basin that doubled

A run of consecutive starting angles reaching one destination is a basin, and its width is a lower bound. Halving the spacing doubled the angles in the widest one and left its width alone, which is what a real basin does and a sampling artefact does not.

emergence · Settling

Named alongside it

The objects these essays reach for when they reach for this one.

Honest limitsAttractorMeasurementClaim testingThe placement ruleNegative resultDivergence angleSettlingStarting angleMeasurement errorSamplingResolution

All concepts