Stems and cones

The front that reads one short

Eleven cells of a fourteen-cell design put the boundary exactly at the larger spiral count. Three put it one offset earlier, and the tempting move is to lower the threshold until all fourteen agree. Measured instead of tuned, the three turn out to be the three cells nearest below their own rung's boundary — and the last offset of a front is weak because it has only just arrived.

Worth reading first: The rate decides the branch · The organ that was taken away · The counts change with radius.

The front result says that removing one organ is felt out to the larger of a stem’s two spiral counts and no further. Put to a design of fourteen cells — seven rises, two branches — it holds at eleven of them exactly. At the other three the run of felt offsets stops one place early: a 4/7 stem whose front reads six, a 7/11 stem whose front reads ten, an 8/13 stem whose front reads twelve.

There is a cheap way to make that go away. The boundary is decided by a threshold — the next organ counts as moved if it moves by more than two and a half degrees — and the offsets in question move it by 1.88°, 2.34° and 1.41°. Lower the threshold to one degree and all fourteen cells agree.

That would be a mistake, and not a subtle one. A threshold chosen so that a claim comes out true is not a measurement of anything, and the resulting statement — the front is always the larger parastichy number — would be unfalsifiable by construction. So the three cells are reported as they are, and this essay is about what they turn out to be.

The same rule, the same rise, two lattices, two frontsHow many organs back a single removal is still felt, on a stem seeded onto the golden lattice and on one seeded onto the Lucas lattice, at seven rises. Everything but the seed is identical at each rise — the rule, the spacing, the heights, the azimuth grid — and the two fronts differ at every one of them. Which branch has the wider front changes hands four times going down the range, so no function of the rise gives the column. The golden branch carries 5/8 at four of these rises, across a factor of two in the rise, and its front is eight at all four.456781011121.621.701.801.8922.102.22rise (falling to the right)how deep the front is, in organs3/53/44/75/87/118/13goldenLucas7 rises · both branches settled to under 0.5°4 reversals
Fig. 1 The design, with the three short cells in it. The Lucas stem at a rise of 0.020 reads six against a count of seven; at 0.008 it reads ten against eleven; the golden stem at 0.006 reads twelve against thirteen.
Which offsets give short hops, at a rise of 0.016The two lowest points are at 5 and 8, and those are the parastichy numbers. Offset 1 is high because a hop of one node is at least the rise, which is what makes a stem easier to count than a disc.00.2000.4000.600102030index offsetmedian hop between node i and node i+m58300 nodes, 34 offsets triedshortest at 5 and 8
Fig. 2 The lattice at the topmost cell of the golden branch’s 5/8 rung, ranked by step length. The two lowest points are close together here in a way they are not further down the rung, and that closeness is what the essay is about.

What the boundary looks like when it is sharp

Start with a cell where nothing is in doubt, because the contrast is the whole argument.

At a rise of 0.010 the golden stem carries 5/8. Removing the organ one place back moves the next one by a large angle; so does two, three, and on out to eight. The displacement at offset eight is 8.91°. At offset nine it is 0.47°, at ten less still, and it stays there out to twelve.

The next organ moves for the last 8, and for no othersOne row per organ removed, counted back from the tip of a stem at a rise of 0.013 whose counted pair is 5 and 8. Removing any of the last 8 moves the next organ by 4.9° to 164.1°; removing an older one moves it by at most 0.70°, which is under the azimuth grid. The boundary is at 8, and 8 is the larger parastichy number — so the experiment counts the spirals without measuring an angle.organ removed, counted back from the tiphow far the next organ moves, in degrees1136.9°286.0°348.3°4164.1°526.2°692.3°7131.0°84.9°— the front ends here90.0°100.7°110.7°120.0°130.7°140.0°150.2°160.2°rise 0.013 · pair 5/8generated from a stated rule, not drawn to look right
Fig. 3 A sharp boundary. Inside the front the displacements are several to a hundred and seventy degrees; outside it they are under half a degree. There is no intermediate region for a threshold to be argued about.

That is a step with a factor of twenty across it, and any threshold between one and eight degrees returns the same answer. The measurement does not depend on the number chosen, which is what a good boundary looks like.

Now the same measurement at a rise of 0.016, on the same branch and the same lattice. The displacement at offset eight is 2.58° — barely over the threshold — and past it, 0.70°. The step is still there and it is much shorter.

Every transition as the rise fallsThe pair climbs 1/2 → 2/3 → 3/5 → 5/8 → 8/13. Consecutive transitions are 0.382, 0.380, 0.382 of the previous rise — 1/φ² is 0.3820.0.4000.6000.8001-2-1.50-1-0.500log₁₀ of the rise between nodes (falling to the right is the plant growing)log₁₀ of the larger parastichy number2/33/55/88/13500 rises, shortest vectors recomputed at eachratio 0.3814 against 1/φ² = 0.3820
Fig. 4 The boundaries a stem crosses as its rise falls. A cell of the design sits somewhere between two of them, and where it sits is the variable this essay adds.

The three short cells are the three newest rungs

The pattern in that is not obvious from the rises and is obvious from the ladder.

A stem’s parastichy pair changes as its rise falls, and the rises at which it changes are computable: they are the rises at which two lattice steps have equal length, solved from the stem’s own settled divergence rather than fitted. The golden branch enters 5/8 at a rise of 0.0182 and leaves it at 0.0069. The Lucas branch enters 4/7 at 0.0249 and leaves it at 0.0095.

Line the design’s cells up against those boundaries and the three short cells are the three that have most recently entered their rung.

The newest member of the front is the weakestFor every cell of the design whose rung boundary is inside the range, how far the next organ moves when the organ exactly as many places back as the larger parastichy number is removed — the offset that arrived when the stem entered this rung — against how far below that boundary the stem sits. Each line is one lattice on one branch. The horizontal line is the threshold that decides whether an offset counts as felt, and the three cells below it are the three whose front reads one offset short. Nothing is a different kind of thing: the boundary is a step everywhere, and near the top of a rung its last stair is shallow.010201.502how far below its rung's own boundary the stem sitsdisplacement at that offset (°)golden 3/5Lucas 4/7golden 5/8Lucas 7/11golden 8/13felt above 2.5°rung boundaries solved, not fittedgenerated from a stated rule, not drawn to look right
Fig. 5 The measurement that explains them. For every cell, how far the next organ moves when the organ exactly as many places back as the larger count is removed, against how far below its own rung’s boundary the stem sits. Each line is one lattice; the horizontal line is the threshold.

The golden 5/8 cells sit at 1.13, 1.39, 1.82 and 2.27 times below their entry rise, and the displacement at offset eight runs 2.58°, 4.92°, 8.91°, 11.95°. The Lucas 4/7 cells sit at 1.19, 1.53, 1.91 and 2.50, and the displacement at offset seven runs 1.88°, 6.33°, 11.25°, 15.47°. The Lucas 7/11 pair gives 2.34° and 5.16°; the golden 8/13 cell, alone in its rung and only 1.14 below its boundary, gives 1.41°.

Every one of those sequences increases, at every step, without exception. The three cells that read short are the three lowest values, and each is the topmost cell of its own rung.

On a rung the response is a run of offsets; near a transition it has a holeOne row per rise, from 0.032 at the top to 0.005 at the bottom, and one column per offset: the organ one place back at the left, 13 places back at the right. A cell is filled where removing that organ moves the next organ by more than 2.5°, and empty where it does not. On a rung the filled cells are a run from one to the larger parastichy number — 5, 8, 13 at the pairs shown on the left. Every rise shown here is in the middle of its rung, so every row is a run and nothing past it is felt — what happens between the rungs is a separate matter.riseorgans back from the tip →run · isolated246810120.0323/550.0135/880.0058/13133 rises · 1536 azimuthsgenerated from a stated rule, not drawn to look right
Fig. 6 The response at three rises well inside their own rungs, where the boundary is unambiguous. Every cell of that table clears the threshold by a wide margin, which is the state the three short cells are being compared with.

The three cells, one at a time

Averages and trends hide the cases they are made of, so here are the three, with what each one actually shows.

The Lucas stem at a rise of 0.020. Its settled divergence is 101.76° and a blind counter returns 4 and 7. The run of felt offsets is one through six; offset seven displaces the next organ by 1.88° and offsets eight through eleven by less. Its rung entry is at a rise of 0.0249, so it sits 1.19 times below the boundary — the second-highest position in the whole design.

The Lucas stem at 0.008. Divergence 99.08°, counted 7 and 11. Offsets one through ten are felt; offset eleven gives 2.34°, which is within a fifteenth of the threshold, and past it the largest anywhere is under half a degree. It entered 7/11 at a rise of 0.0095 and sits 1.19 below it.

The golden stem at 0.006. Divergence 137.93°, counted 8 and 13. Offsets one through twelve are felt, offset thirteen gives 1.41°, and it entered 8/13 at 0.0069, sitting 1.14 below. It is the newest rung in the design and it gives the smallest displacement in the design.

One rise, two seeds — the response of eachHow far the next organ moves when the organ a given number of places back is removed, at a rise of 0.008, on a stem seeded onto the golden lattice and on one seeded onto the Lucas lattice. The shading is the size of the displacement and the vertical line is where the run of felt offsets ends: 8 on the golden stem, whose lattice is 5/8, and 10 on the Lucas stem, whose lattice is 7/11. Nothing else differs between the two runs. Everything that changes with the rise — the spacing, the heights, the size of the neighbourhood the rule sums over — is the same in both rows.123456789101112golden5/8, front 8123456789101112131415Lucas7/11, front 10organ removed, places back from the tiprise 0.008 · same rule, same grid, same heightsfronts 8 and 10
Fig. 7 One of the three in its own row, beside the branch that is well inside its rung. The golden row’s run ends squarely on eight; the Lucas row’s ends at ten with an eleventh cell that is faintly shaded rather than blank.

The ordering by position in the rung is exact: 1.14, 1.19, 1.19 for the three short cells, against 1.13 for the one cell that clears the threshold by a fourteenth of a degree and 1.39 and upwards for everything that clears it comfortably. There is one apparent inversion — the golden 5/8 cell at 1.13 sits lower in its rung than any short cell and reads correctly — and it is the useful part of the data rather than an embarrassment: at 2.58° it is on the threshold, and it says the effect is a continuous shortening of the last stair rather than a switch that flips at a particular depth.

What a divergence picked at random gives, at a rise of 0.016Fibonacci pairs take 20.5% of the circle at this rise, and the share falls as the rise does. The claim that Fibonacci counts are what nature "prefers" needs the preference to come from somewhere, and it is not from the geometry being generous.Fibonacci20.5%Lucas3.8%whorled31.4%other44.2%25 distinct pairs over 1200 divergencesrise 0.016Fibonacci 20.5%
Fig. 8 Every pair available at the rise of the topmost cell, ranked by how short its steps are. Near a boundary the second and third entries are nearly tied, which is the same fact the essay reads in the response.

Why the newest offset is the weakest

The reason is what a rung boundary is.

At a boundary, two candidate second families have equal step length. Just below it, the new family’s steps are barely shorter than the old family’s, and the new count has only just become the larger of the pair. So an organ exactly that many places back is, at that rise, only marginally a member of the neighbourhood the rule is sensitive to. Further down the rung its steps shorten relative to everything else and it becomes solidly one.

The plane of stems: divergence across, rise upEach shade is one parastichy pair. The marked points are the lattices where three families are equally short — the forks — and the Fibonacci ones run up the middle towards 137.51°.-2-1.50-1-0.500100120140160180divergence angle (°)log₁₀ of the rise between nodes1,2,32,3,560 × 150 lattices, each solved586 runs drawn
Fig. 9 Where the boundaries come from. The branches meet at rises where two lattice steps have equal length; a stem just below such a meeting is one whose newest family has only just won.

So the front does not have a hard edge that sometimes slips. It has an edge whose last stair varies in height with position in the rung, and near the top of a rung that stair is under two degrees.

This also predicts something the design did not set out to test and confirms it in passing: the second-newest offset should be stronger, and it is. At the golden 8/13 cell — the shortest-lived rung in the design — offset thirteen gives 1.41° and offset twelve gives 24.6°, which is why the front reads twelve rather than eleven.

What the threshold is, and why it is not moved

It is worth stating what the threshold is set from, since the argument above depends on not adjusting it.

Two and a half degrees is a tenth of the local spacing between organs at the rise the ablation work is centred on, and four steps of the azimuth grid the runs are computed on. It was fixed before any of this was measured, on the grounds that inside the front displacements run from a few degrees to a hundred and seventy and outside it they run under half a degree — so anything between half a degree and two degrees gives the same table at the cells where the boundary is sharp.

On a rung the response is a run of offsets; near a transition it has a holeOne row per rise, from 0.032 at the top to 0.005 at the bottom, and one column per offset: the organ one place back at the left, 14 places back at the right. A cell is filled where removing that organ moves the next organ by more than 2.5°, and empty where it does not. On a rung the filled cells are a run from one to the larger parastichy number — 5, 8, 13 at the pairs shown on the left. Every rise shown here is in the middle of its rung, so every row is a run and nothing past it is felt — what happens between the rungs is a separate matter.riseorgans back from the tip →run · isolated24681012140.0323/550.0135/880.0058/13133 rises · 1536 azimuthsgenerated from a stated rule, not drawn to look right
Fig. 10 Why the number was chosen where it was: at the rises the single-organ result was established on, the gap between felt and unfelt spans two orders of magnitude and the threshold has a wide empty band to sit in.

At the three short cells that empty band is narrow, and the honest description is that the measurement has become threshold-dependent there. Reporting eleven of fourteen with the other three named, their displacements printed and their position in the rung explained is what that description looks like. Reporting fourteen of fourteen at a threshold of one degree would be reporting the threshold.

The block a wrecked stem settles into is the count it was cut fromA stem is cut in the middle of its front and followed for 300 organs. It does not come back to its lattice; what it does instead is repeat a fixed sequence of divergences exactly, to the resolution of the azimuth grid. Each row shows that sequence twice over, one bar per organ, drawn to the same scale. At the 5/8 rung the sequence is five angles long and advances -31.4° per block. Every block is the smaller number of the pair that was cut, so the second attractor carries the first one's count — and every precession is one part in twice the block, which makes each of these a two-jugate arrangement with 11 rows.5/8 rungblock of 5-31.4° a blockand again209.7° on average300 organs after the cut · 1536 azimuthsgenerated from a stated rule, not drawn to look right
Fig. 11 One of the other consequences of being near a boundary: at rises where the runner-up is close, a removal at the wrong offset is not repaired at all.

The same shape, elsewhere in this collection

A boundary that is sharp in the middle of a rung and soft at its top is not a new kind of object here, and the earlier cases are worth putting beside it.

The response of a stem to a removal is an interval of offsets only in the middle of a rung. Near a transition it acquires a hole: a stretch of quiet, and then one isolated offset well outside the front that is felt strongly. That was measured on the golden branch and its position was pinned to the count coming in at the next rung, less one.

The blind counter has the same character. Given an ideal lattice in the middle of a rung it returns a pair with a wide margin — the third-shortest step is much longer than the second. Near a boundary the second and third are nearly tied, and the counter’s answer, while still correct, is one a tenth of a degree of noise could change.

Two answers 138° apart, and one organ holding the second one upThe repulsion the rule minimises, around the circumference of a stem at a rise of 0.008, at the height the next organ will sit at. It has two low points 138.3° apart: the slot the next organ takes, and the slot the organ after it will take. The runner-up is 13.6% higher. The organ 13 places back carries 14.6% of the energy at the winning slot and twelve places back carries 16.3% at the runner-up — and that is more than the gap, so taking that organ away makes the runner-up win and the next organ appears a whole divergence away. Neither guard is a contact of the organ being placed; twelve is one place inside the larger number of the pair this stem is climbing towards.the slot it takesthe slot after next, 14% higherazimuth around the stemrepulsion around the circumference13 back holds the first, twelve back holds the secondrise 0.008 · pair 5/8 · climbing to 8/13generated from a stated rule, not drawn to look right
Fig. 12 The structure underneath both: near a transition the runner-up is close behind the winner, so every quantity that depends on which family is second becomes delicate at the same rises.

So this essay’s finding is the third instance of one thing. Every property of a lattice that depends on which family is second degrades near the rise where two families are tied, and the front is such a property, because the front’s depth is the larger of the two counts. Reading that as a defect of the front measurement would be reading a property of lattices as a property of an instrument.

A consequence for anybody doing this on a plant

The result has a use, and this essay changes what it is worth in a specific way.

The attractive thing about the front is that it turns a spiral count into a sequence of yes-or-no answers, which is a far cruder measurement than counting spirals on a photograph. This says the crudeness has a limit that depends on where the specimen sits on its own ladder. On a stem well inside a rung, the last offset of the front displaces the next organ by ten degrees or more and any competent observation catches it. On a stem that has just changed rung, the last offset displaces it by one or two degrees, and an observer will systematically read the front one short.

Every transition as the rise fallsThe pair climbs 1/2 → 2/3 → 3/5 → 5/8 → 8/13. Consecutive transitions are 0.381, 0.382, 0.382 of the previous rise — 1/φ² is 0.3820.0.4000.6000.8001-2-1.50-1-0.500log₁₀ of the rise between nodes (falling to the right is the plant growing)log₁₀ of the larger parastichy number2/33/55/88/13500 rises, shortest vectors recomputed at eachratio 0.3816 against 1/φ² = 0.3820
Fig. 13 Where a specimen sits on its ladder is not something an experimenter chooses. A stem grows down this axis, so a plant caught shortly after a transition is a plant whose front will read short.

That is a bias rather than noise: it always errs in the same direction, and it errs on exactly the specimens that have most recently changed their counts. A census of ablation experiments that did not record where each specimen was on its ladder would under-report the front, by one, on a subset it could not identify afterwards.

The repair is cheap and it is the same one this collection keeps arriving at: record the counts and the geometry. A specimen whose pair has just changed is identifiable — its two families have nearly equal step lengths, which is measurable on the same photograph the counts come from.

What the positions say, and what the angles sayEach row is one stem at one rise. The left column is the parastichy pair counted from the coordinates; the right is the single number read out of the divergence angles alone, over 4 runs. On the Lucas ladder — 3/4, 4/7, 7/11 — the readout returns the smaller number too, so it is reading the lattice rather than Fibonacci. The last row is the one that matters: at a rise of 0.05 the positions give an unarguable 2/3 and the angles give 4, 23, 12, 2 — all five wrong, and all five refused.counted from the pointsread from the anglesgolden, rise 0.0323 / 534/4 clear · peak 0.73golden, rise 0.0135 / 854/4 clear · peak 0.61golden, rise 0.0058 / 1384/4 clear · peak 0.78Lucas, rise 0.0323 / 434/4 clear · peak 0.55Lucas, rise 0.024 / 744/4 clear · peak 0.46Lucas, rise 0.0087 / 1174/4 clear · peak 0.60golden, rise 0.052 / 34, 23, 12, 2refused — peak 0.12 under 0.344 runs per rise · the readout sees a list of angles and nothing elsethe refusal is the gate working
Fig. 14 The two ways a lattice can be read — from the sequence of angles and from the positions — and what each costs. A yes-or-no answer about a removal is far cheaper than either, which is the whole appeal of the front.

What a soft edge does to the yes-or-no reading

The front’s appeal is that it converts a count into a run of answers to a much easier question. It is worth working out what a soft last stair costs that conversion, because the answer is not “a bit of noise”.

Suppose an observer removes an organ at each offset in turn on a set of clones and records whether the next organ moved. Inside the front the displacements are tens of degrees and the answer is unambiguous however the observation is made. At the last offset of a stem near the top of its rung the displacement is one to two degrees, which is comparable to the divergence scatter this collection measures in its own noisy runs and comparable to what a protractor on a real apex would resolve.

So the failure mode is not a wrong count with a wide error bar. It is a count that comes out exactly one less, confidently, with every other offset behaving perfectly. An experimenter would have no internal signal that anything had gone wrong: the run would be consecutive, the region past it quiet, the boundary clean.

The measurement is limited by the protractor, not by the plantThe peak falls as the reading error grows, and it falls by an arithmetic factor with nothing fitted: a position error enters two consecutive divergences with opposite signs, adding variance at every lag while the pattern's signal sits at one. At a quarter of a degree the readout is right on all 5 runs; at half a degree on 2; at a degree on 1. Below the dashed floor the peak is the largest of thirty noisy numbers rather than a measurement.00.2000.4000.6000.80000.50011.502reading error on each organ's position, in degreesheight of the peak at the parastichy numberwhat noise alone givesthe threshold a reading must clear5/5 right5/5 right2/5 right1/5 right1/5 rightpredictedrise 0.008 · 5 runs · pattern scatter 0.75°peak × σ²/(σ² + 2ε²), nothing fitted
Fig. 15 The general form of the problem this collection keeps meeting: what an instrument can resolve decides which of two nearby answers it returns, and the returned answer carries no trace of the decision.

That is the reason to state the three cells rather than tune them away. A known bias of exactly one, on an identifiable subset of specimens, is something a census can correct for. An unstated one is a census that reports the wrong integer.

What this does not say

It does not say the three cells refute the front result. At each of them the run of felt offsets ends one place early and everything past it is quiet, so the shape of the result is intact; what varies is the height of the last stair. A cell whose front read four short, or whose felt offsets were scattered rather than consecutive, would be a different matter and none does.

It does not say the strengthening is linear, or that it has been characterised. Four points per lattice on two lattices, plus two shorter series, is enough to say the ordering is always the same and not enough to fit anything. The claim is monotonicity and nothing more.

It does not say the threshold is arbitrary. It is set from the spacing and from the grid, and at eleven of fourteen cells it could be moved by a factor of five with no effect. The three cells are where that stops being true.

And it does not explain the two cells that are not in the pattern. The Lucas stem at a rise of 0.024 carries 3/4, which the ideal ladder says it should already have left; it is holding a pair past its own boundary, so it has no position in its rung to measure and it is excluded from the comparison rather than fitted into it. That hysteresis is real and is another thread’s subject.

The check that would refuse it

Two assertions run with the figure, and the second is the interesting one.

The first is that within each lattice appearing at two or more rises, the displacement at the offset equal to the larger parastichy number grows at every step down the rung. That is four sequences with no exceptions allowed, and a single inversion anywhere stops the collection being built. It is the assertion that would fail if the pattern were a coincidence of two cells.

The second is that every cell whose front reads short is one whose newest offset falls under the threshold — which sounds circular and is not. The front reading short means the run of consecutive felt offsets ends early, and it would also happen if some offset in the middle of the run were quiet, or if the offsets past the boundary were noisy enough to make the run ambiguous. The assertion says the short reading is always attributable to that one offset, and it would fail if a short cell were short for any other reason.

There is a third guard on the cells that are excluded. A stem carrying a pair the ideal ladder says it has already left has no measurable position in its rung, and rather than fitting one it is dropped from the comparison and reported as lagging. That is one cell of fourteen, and saying so is cheaper than the alternative, which is a design that quietly contains a specimen it cannot place.

Shares its objects with

Essays that name at least two of the same things, and that neither author linked.

  • The response with a hole in it — both name ablation, artefact, discretisation, honest limits, identifiability, ladder, measurement, parastichy pair, rise, rung, transitions
  • A cut of two organs — both name ablation, artefact, discretisation, honest limits, ladder, measurement, parastichy pair, rise, rung
  • The organ that guards the second slot — both name ablation, artefact, discretisation, falsifiability, honest limits, measurement, parastichy pair, rise, transitions
  • The rung was not the instrument — both name artefact, discretisation, honest limits, identifiability, ladder, measurement, parastichy pair, rise, rung
  • The ratio was the floor of a curve — both name artefact, honest limits, ladder, measurement, parastichy pair, rise, rung, transitions
  • What a sample grid decides — both name artefact, discretisation, falsifiability, honest limits, identifiability, measurement, parastichy pair, rise

Named objects

A flat tag is an object no other essay names yet.

AblationArtefactBranchDiscretisationFalsifiabilityHonest limitsIdentifiabilityLadderMatched designMeasurementParastichy pairRiseRungToleranceTransitions