Phyllotaxis
the spirals are counted, not admired
Concept

Tolerance — where it appears

The margin inside which two computed numbers are called equal, stated before the comparison rather than after it. Every tolerance in this collection is set from something measurable — a grid step, a sampling error, a spread between seeds.

Named by 34 essays across 6 fields — each of them below, with the objects they name alongside it.

Three disturbances, three places to get in. The rule reads its neighbours, builds a profile of the energy at every azimuth, takes the least of it, and records a position. field noise enters at the profile; jostle noise enters at the neighbours; placement noise enters at the record. Two of the three are upstream of the choice and can change which minimum is taken; the third is downstream and never can.

The noise that arrives through the neighbours

The two kinds of noise this site had were idealisations that bracket the rule's choice. The realistic disturbance is neither: a primordium is placed exactly, and then the organ grows, so by the time the next one forms its neighbours have moved. That is a third kind, and it is invisible in every measurement a plant offers.

mechanism · Noise entry A lattice survives about 1.8° of scatter, whichever way the noise arrives. The largest divergence scatter at which a run is still a lattice, from each kind of noise at the largest amplitude that leaves one. The two routes share no code below the placement rule: one displaces the node after the choice, the other perturbs the energy the choice is made over. They agree to 0.33°.

Two degrees of scatter

A lattice tolerates about two degrees of wander in its divergence angle, and two kinds of noise sharing no code agree on the number to within a third of a degree. It is not a constant: carried finer, the same stem survives 0.8°, and the tolerance tracks the band of angles that produce its pair at all.

emergence · Scatter tolerance What the sequence sees that the scatter cannot. Each point is an ensemble at one amplitude, placed at the scatter it produces. A stem at three quarters of a degree of scatter has a lag-one correlation near zero if its noise arrived after the primordium was placed, and near 0.7 if it arrived before — and no measurement of the scatter can tell those apart. The separation closes above about a degree, because what the other two kinds preserve is the correlation of a lattice.

What a quiet plant is worth

Almost every measurement gets easier as the effect gets larger. This one gets harder — a stem's divergence sequence stops carrying information about its noise at precisely the scatter where the noise becomes obvious. The specimens worth measuring are the ones that look least interesting.

wrong · Survey spec Four fractions of 34, one width. The half-width of the second-moment dip at four divergence angles with the same denominator, on heads of 791 organs — 23 in each of 34 rows. 13/34 is the Fibonacci convergent, the fraction every earlier measurement of this law was made at and the one whose neighbourhood is emptiest; 9/34, 15/34, 11/34 are not. The four agree within a factor of 1.14, well inside the factor of two the law is stated to, so the width is a function of the denominator and not of how well the fraction approximates its neighbours. What is left over is ordered by the note beside each bar: the more crowded the neighbourhood, the wider the dip comes out, which is the direction a neighbour's own shoulder would push it.

A width read off a staircase

Two fractions of the fourteen measured return a dip width that moves by a factor of two when the head size changes, where the others hold to three per cent. The cause is not their neighbourhood. It is that the disorder statistic changes only when the tessellation changes, so the curve a half-width is read off is a staircase, and a width narrower than the tread cannot be read at all.

tissue · Second statistic What insisting on a fork angle costs: 43.3 degrees for one per cent of the network. The three ends are held where the optimum wants them, the branch point is moved everywhere inside them, and the cheapest network at each total angle is kept. The optimum sits at 74.93°. Everything within one per cent of the least cost runs 55.9° to 99.2° — a span of 43.3°, which read back as exponents covers 2.44 to 5.34 — and within a tenth of a per cent it still runs 13.6°. The prediction is steep in the exponent and the cost is nearly flat in the angle; they are the same curve read along its two axes.

An optimum too flat to reach

One per cent of a branching network's cost buys forty-three degrees of fork angle, covering exponents from 2.44 to 5.34, while the angle the theory predicts moves only fourteen and a half degrees across every daughter ratio there is. The prediction is steep and the cost is flat, and those are the same curve read along its two axes.

branching · Fork angle The two trees this site draws, at 30° and 32° to a side, against the cost's 37.47° and 37.47°. Two trees of 63 segments each, 5 generations deep and 31 junctions apiece, with every junction's radii taken from r₀³ = r₁³ + r₂³ exactly and every junction's angle taken from a constant. Read as an exponent through cos(θ/2) = 2^(2/p − 1), the drawn angles say 2.5237 and 2.6239, in pictures whose widths are built at exactly 3. The cost that fixed those widths wants 37.47° and 37.47° at this daughter ratio, 74.93° in total, and the misses cost 0.573% and 0.292% of the network — which is why a fixed angle can sit in a figure about a minimisation and never look wrong.

The trees drawn at no angle

Two branching figures in these essays set every junction's radii from the cube law exactly and every junction's angle from a constant nobody derived. Read as exponents the drawn angles say 2.52 and 2.62, in pictures whose widths say exactly three — and at a lopsided fork the drawing puts a daughter thirty-four degrees from where the same cost puts it.

branching · Fork angle What the sequence sees that the scatter cannot. Each point is an ensemble at one amplitude, placed at the scatter it produces. A stem at three quarters of a degree of scatter has a lag-one correlation near zero if its noise arrived after the primordium was placed, and near 0.7 if it arrived before — and no measurement of the scatter can tell those apart. The separation closes above about a degree, because what the other two kinds preserve is the correlation of a lattice.

What one angle says about the next

A tenth of a degree of placement noise moves a stem's divergence scatter from 0.50° to 0.62°, which nobody would report. It takes the correlation between consecutive angles from 0.54 to below zero. The other two kinds of noise, at scatters where no measurement can separate them, leave it at 0.6.

cylinder · Sequence Both statistics, on the same stems, at a rise of 0.005. Five seeded stems at each disturbance, held at a fixed rise. Bars are how many returned the pair the position counter finds; open portions are refusals. The pair comes out from 0.1 to 0.25, and across that whole range the lag-one correlation of the same sequences is -0.33, -0.58, -0.59 — decisive, negative and flat. There is no trade between the two: one stem supplies both. Below the window the sequence has locked onto the sampling grid and is a cycle rather than a sample; above it there is no lattice left, at 117° of scatter.

What a refusal does not say

The readout can decline for four different reasons — too quiet, too disturbed, too fast, or a window in the wrong place — and a stem that returns nothing does not say which. That is the third time this thread has failed to close the mixture problem, and the first time the failure has a shape.

wrong · Survey spec Through the rule, the drift survives and the inheritance still does not. How much of a divergence sequence's variance survives being averaged over blocks, on stems the rule grew. The vertical quantity is B² times the variance of the block means divided by the variance of the sequence, which is one at every block size for independent errors — the arithmetic is normalised for a differenced stream, since a divergence is the difference of two organs' errors. A line that climbs is a sequence with power at frequencies below one per block. The disturbances that remember the last error climb to 46 at a block of 64. The ones inherited between touching organs do not climb at all — 1.51 and 1.90 at the same block — although their own deviates carry ×— and ×— an independent stream's variance in exactly this statistic. What they do instead is dig a hole: at block sizes of 8 and 13, which are the offsets they couple at, the statistic falls to 0.06 and 0.12.

What the rule does to a drift

A placement rule was supposed to leave no slow wander in a divergence sequence, because its errors are corrections rather than inheritances. Driven by a disturbance that drifts, it leaves a larger one than a lattice with no rule in it at all — while cutting the per-organ scatter by more than half. The rule removes what is relative between neighbours, and a drift is not.

mechanism · Noise colour Both ends of the window are silent, and a ruler tells them apart. The scatter recorded on stems at 400 nodes per rung, against the disturbance that produced it, with the stems that returned no reading at all marked as open. Silence at the quiet end comes with a scatter of 0.38 and 0.44°, which any botanist would call an orderly plant; silence at the disturbed end comes with 56°, which nobody would call a pattern. The two refusals look identical in the instrument's output and are three orders of magnitude apart in a quantity measured with a protractor.

A refusal with a reason

Three note left with the work running have recorded that a refusal has four causes and the sequence separates none of them. With a second window and a protractor, three are separated: silence at 0.38° of scatter is a quiet plant, silence at 56° is a disorderly one, and agreement certifies the rate. The fourth survives, and so does a worse discovery — agreement is not correctness.

wrong · Two windows Which lattices survive a fifth of a degree of noise. The share of runs that still have a lattice. The prediction was that a cut-off at this range would be as fragile as the truncation it replaces; it is not. Stating the neighbourhood as a function of distance did not merely make the old result honest — 100% of runs survive against 33%, at a scatter an inverse-cube rule cannot be told from.

The fragility belonged to the window

A pattern that exists only because the rule cannot see far was expected to be held together by that cut, and to fall over when nudged. It does — while the cut is a loop bound. Written down as a falloff at the same range, the same rule keeps every run under the same nudge, at a scatter an inverse-cube rule cannot be told from.

emergence · Noise amplitude Both statistics, on the same stems, at a rise of 0.005. Five seeded stems at each disturbance, held at a fixed rise. Bars are how many returned the pair the position counter finds; open portions are refusals. The pair comes out from 0.1 to 0.25, and across that whole range the lag-one correlation of the same sequences is -0.33, -0.58, -0.59 — decisive, negative and flat. There is no trade between the two: one stem supplies both. Below the window the sequence has locked onto the sampling grid and is a cycle rather than a sample; above it there is no lattice left, at 116° of scatter.

Two readings from one stem

Three note left with the work in a row have recorded that the two statistics of a divergence sequence want opposite plants — one quiet, one disturbed. Measured on the same stems they do not. The conflict was in the interpretation of a sign, and the window in which both are readable is wide.

cylinder · Sequence Placements that went to a different minimum, per thousand. The rule's own counterfactual, run beside it: where would this node have gone with the noise taken away and everything else left alone? Placement noise displaces the node after the argmin, so the answer is always "here" — 0.2°, 0.4°, 0.8° all give zero. A jostle and a field perturbation are upstream of the choice and change one or two placements in a thousand while the lattice is still intact.

Which minimum was chosen

The rule takes an argmin, so there are two completely different things noise can do to it: move the answer, or move the question. One of them can change what is chosen and the other cannot, ever — and the difference is exactly zero against one or two placements in a thousand, at amplitudes where every other measurement says the two are identical.

emergence · Noise entry Four ways to count the rule's neighbourhood, and one ordering. How many organs the placement rule is looking at, as its falloff exponent is swept from 1.5 to 6, by four different definitions. Counting the organs that carry half the profile gives 33, 12, 3, 2, 1. Counting the organs that carry nine tenths gives 182, 120, 30, 8, 3. Counting the organs that carry ninety-nine hundredths gives 284, 263, 149, 54, 9. Counting the organs that carry the weighted mean lag gives 68, 45, 21, 14, 11. They disagree about the size of the neighbourhood by a factor of 10 — the widest moves by 61 times across the sweep and the narrowest by 6.1 — and every one of them falls as the exponent rises. So no choice among them changes which rule is the deeper, and no redefinition can turn a result about the depth around.

Four ways to count a neighbourhood

How deep does the placement rule look? Counting the organs that carry nine tenths of its profile gives 182 down to 3 as the falloff steepens. Counting the ones that carry half gives 33 down to 1. The weighted mean lag gives 68 down to 11. The four disagree by an order of magnitude about the size and agree exactly about the order.

mechanism · Noise colour Where each kind's lattice gives way. The largest amplitude at which every run still has a lattice, and the scatter it produces there. The amplitudes are incomparable — field 0.015 (fraction of the barrier), jostle 1 (degrees of azimuth), placement 0.8 (degrees of azimuth) — and the scatters agree to 19%. The boundary belongs to the pattern rather than to the disturbance: a lattice fails at about a degree and a half of scatter, and which of three mechanisms produced it does not move where.

The boundary belongs to the pattern

Three kinds of noise, in three incommensurable units, destroy a lattice at the same place — about a degree and a half of divergence scatter. The earlier work measured that of two kinds and called it a scale rather than a constant. With a third it looks less like a coincidence and more like a property of what a lattice is.

emergence · Scatter tolerance At 384 azimuths the ratio is 0.62; converged it is 0.82. The comb ratio and the recorded divergence scatter at a rise of 0.005, against how finely the rule samples the circle when it takes its minimum. At 384 azimuths — the grid every flat run in these essays uses, and the grid that earlier work's 0.65 was measured on — the step is 0.94°, which is larger than the 0.25° disturbance the stems carry. The quantisation is white noise, it dilutes both combs, and it does not dilute them equally. The ratio settles at 0.82 from 1152 azimuths up, and the scatter loses 0.19° that belonged to the grid rather than to the stem.

The grid was in the number

The rule places each organ at the least of a profile sampled at a fixed number of azimuths, and every flat run in these essays samples 384 of them — a step of 0.94°, against a disturbance of a quarter of a degree. The quantisation is the larger of the two, it is white, and it moves the discriminator from 0.79 to 0.62. The convergence study this collection had asked for and never done, in the place it turned out to matter.

cylinder · Instrument ceiling The damage is the sharing; the forgery is the history. Three disturbances of the same size, measured four ways. The two left columns are stems grown by the placement rule and jostled at 0.25° per organ: a disturbance shared between the contact neighbours scatters the lattice by 0.71° against white noise's 0.57°, and one inherited from them — the same sharing, passed on again at every organ — by 0.97°. The two right columns are kinematic lattices with no rule in them at all, where the whole question is what a disturbance can manufacture. The inherited one returns the pair on 8 seeds of 8 with a main comb of 0.205 against a band of 0.073; the shared one, at the same coupling and the same scatter, returns it on 1 and makes a comb of 0.099, which is the band. So sharing an error with the organs you touch does the damage, and only passing it on and on forges the evidence.

What the sharing costs a lattice

A disturbance inherited from the contact neighbours destroys a stem's lattice at half the displacement independent noise needs, and it moves the comb ratio a fifth of the way to a forgery's. Take the inheritance out and keep the sharing, and the damage stays and most of the ratio shift goes — so the two effects have different causes.

cylinder · Noise transport A cut eight back is never undone. The divergences of a stem whose organ eight places back was removed, against the same stem uncut. It never returns. What it settles into repeats exactly every 8 organs — 47°, 96°, 137°, 273°, 138°, 271°, 230°, 272° — and holds that cycle for the whole 300-organ run, with a mean of 186° and a spread of 83°. A rule that corrects a displacement does not correct a deletion.

A rule that cannot heal a hole

The placement rule corrects itself against a displacement — that is what the lag-one correlation of −0.6 has been saying since it was measured. It does not correct itself against a deletion. Which organ is removed decides whether the stem is back on its lattice in twenty-four organs or never, and the boundary between the two is sharp, reproducible and in the middle of the front.

emergence · Ablation A period of 8, and the two classes that are not with the rest. The same wrecked stem, folded on the lag it kept: one row per residue class, each drawn at the mean displacement of its own organs against the level the rest of them share. The bar through each row is the spread inside that class, and the widest of them is 0.87° — so within a class the displacement is a constant. six classes sit at the common level. The two that do not sit at 134.3° and -134.2°, equal and opposite to within 0.0 per cent, and they are neighbouring residues. The stem's own divergence is 137.44°, so an exception is one organ's step.

One level and two exceptions

Inside a wrecked stem's period most residue classes sit at one level and a couple do not. On seventeen of the thirty cuts the exceptions are exactly two, equal and opposite to within five per cent — and on all seventeen they are neighbouring residues, which was not looked for.

mechanism · Damage shape The exceptional pair, measured in divergences. One row per wrecked cut whose profile has exactly one pair of exceptional classes, drawn at the size of that pair divided by the stem's own settled divergence. Every row sits between 0.882 and 1.076, so the two chains that came apart moved by one organ's step rather than by two or by half of one. The residual is not scatter: rows are grouped by the lag the stem kept, and every lag sits wholly above the line or wholly below it. Why a surviving 5 or 7 overshoots and a surviving 4 or 8 falls short is not answered here.

A step of one organ

The balanced pair inside a wrecked stem's period measures 88.0° to 147.2° against divergences of 99.1° to 138.0° — one organ's step, to within twelve per cent on every row. The residual is not scatter: every stem keeping a 5 or a 7 overshoots and every stem keeping a 4 or an 8 falls short.

mechanism · Damage shape What a lattice survives depends on the colour of the disturbance, sixfold. five stems for each of seven disturbances at each of six displacements, every stream normalised by its own measured spread so that a displacement of half a degree is half a degree in every row. A filled mark is a stem that still has a lattice — a divergence scatter under 2° — and an open one is a stem that does not. Independent errors survive to 0.5°; errors that remember the last one to 1.5°; errors inherited from the contact neighbours only to 0.25°. The number beside each row is the scatter a protractor would record where the lattice is standing, and it is the quantity that explains the table: what destroys a lattice is not how far an organ moves, but how far it moves relative to the organs it is placed against.

A disturbance the organs share

This collection has put three kinds of noise into the placement rule and found the lattice fails at about the same recorded scatter whichever kind it was. None of them asked what happens when the displacements are correlated between organs. At equal displacement per organ, a lattice survives three times as much of a disturbance the organs share — and what a protractor records is the part they do not.

emergence · Noise amplitude What a lattice survives depends on the colour of the disturbance, sixfold. five stems for each of seven disturbances at each of six displacements, every stream normalised by its own measured spread so that a displacement of half a degree is half a degree in every row. A filled mark is a stem that still has a lattice — a divergence scatter under 2° — and an open one is a stem that does not. Independent errors survive to 0.5°; errors that remember the last one to 1.5°; errors inherited from the contact neighbours only to 0.25°. The number beside each row is the scatter a protractor would record where the lattice is standing, and it is the quantity that explains the table: what destroys a lattice is not how far an organ moves, but how far it moves relative to the organs it is placed against.

The disturbance that travels

If a lattice survives three times the displacement when the organs share it, then a disturbance passed between the organs that actually touch should be the gentlest of all — it is correlated at exactly the offsets the rule places against. It is the harshest. Half the displacement destroys what independent noise leaves standing, and the reason separates two things that had been one.

emergence · Noise colour The newest member of the front is the weakest. For every cell of the design whose rung boundary is inside the range, how far the next organ moves when the organ exactly as many places back as the larger parastichy number is removed — the offset that arrived when the stem entered this rung — against how far below that boundary the stem sits. Each line is one lattice on one branch. The horizontal line is the threshold that decides whether an offset counts as felt, and the three cells below it are the three whose front reads one offset short. Nothing is a different kind of thing: the boundary is a step everywhere, and near the top of a rung its last stair is shallow.

The front that reads one short

Eleven cells of a fourteen-cell design put the boundary exactly at the larger spiral count. Three put it one offset earlier, and the tempting move is to lower the threshold until all fourteen agree. Measured instead of tuned, the three turn out to be the three cells nearest below their own rung's boundary — and the last offset of a front is weak because it has only just arrived.

cylinder · Branch choice Everywhere a cut of one to five organs can send a 5/8 stem. Every settled divergence reached by any arrangement of up to five organs removed from a stem at the 5/8 rung, on one axis. There are six of them and no more. three are slips of the lattice the stem was cut from: each keeps the lag-5 family intact and sits a whole number of turns of it from the next, which is the ladder marked below the axis with rungs 72.0 degrees apart. The other three keep no lag at all and sit near a fraction of a turn, marked above: 175.0 degrees near 1 of 2 turns, 190.0 degrees near 1 of 2 turns, 235.0 degrees near 2 of 3 turns. A stem that is cut either slides along the ladder it was on or leaves it for a lattice with files in it.

A file has to close

The three destinations counted with a shared factor sit near a half turn, a half turn and two thirds. Measuring how near is the trap: by distance from the fraction, the golden angle is closer to two fifths than two of them are to anything, and would be reported as having five files it does not have.

cylinder · Dose

The shallower front turns over

If reversing a stem means rearranging its whole front, then a stem with a shallow front should reverse more often. Measured across three rungs and four hundred and seventy-three cuts: 6.8 per cent at a front of three, 4.7 at five, and none at all at eight — where the nearest approach is two tenths of a degree away and stays there.

cylinder · Coarse rung

Twice the run

Five wrecked cuts never reached a pattern inside three hundred organs and one reached it at three hundred and three, which is a number asking to be checked. Run every cut in the census twice as far and three of the thirty change their answer.

mechanism · Run length

Not the shorter of the two

If a damaged stem keeps one contact family standing, the obvious guess is that it keeps the nearer one. Across thirty wrecked offsets that is true twelve times and false seventeen, and on one lattice the two steps differ by a quarter of a per cent — where the words shorter and longer are doing no work at all.

emergence · Ablation

An onset at the end of the run

One cut reported that its pattern began 299 organs into a 300-organ run. Given twice the room it reports no pattern at all. The reading was the run stopping, not the disturbance ending, and the definition guarantees one at the last organ of every run.

mechanism · Run length

Three rows change sides

Twenty-five of thirty wrecked cuts have a periodic displacement profile over three hundred organs and twenty-six do over six hundred. The count barely moves and the membership does: two rows join, one leaves, and the gap the threshold sits in narrows from 1.69 to 1.27.

mechanism · Run length

Four crossings nobody visited

Six rungs of the ladder carry a handover and two of them had a band built on them. The other four are here: 70, 126, 16 and 124 rises wide, found by sweeping at a ratio rather than at a fixed step in the rise, which is why the fine ones had been stepped over.

cylinder · Rung interior

A window nobody aligned

Every reading this thread takes of a wrecked stem is taken inside a window, and there are three of them: a run of three hundred organs, a window of a hundred and twenty at its top, and a search window of fifteen to thirty-nine. None was aligned to anything, and one of them turned out to decide its own answers.

mechanism · Run length

A band that moves nothing

One of the six bands holds the counted pair, holds the divergence, and does not move the ordering: its two contact steps stay within four parts in a thousand of each other across the whole of it, so the ordering changes hands three times and neither end has one worth the name.

cylinder · Rung interior

How wide a band should be

A band ends where the divergence has slid a twentieth of a degree, so its width should follow from how fast the divergence slides. Predicted from the rung's slope that is wrong by factors of 0.20 to 5.92; predicted from a stationary point it is 0.41 to 1.08, and the outlier is the rung that has no stationary point.

cylinder · Rung interior

A list that was a rounding

Three destinations only a steep falloff reaches, read at two grid steps. Thirteen, read at a tenth of a degree. And four arrangements, read by what the counter returns rather than by the angle — a different four, with one the angle reading hides.

emergence · Off-ladder

Named alongside it

The objects these essays reach for when they reach for this one.

MeasurementDivergence angleHonest limitsNoiseEnsembleNegative resultArtefactThe placement ruleParastichy pairSummary statisticAblationAutocorrelation

All concepts