Transitions — where it appears
Counting the spirals
Almost every claim about phyllotaxis is a claim about how many spirals run through a pattern, and the count is almost never done. It can be done from the points alone, by machinery that is never told what angle built them — and then a count of 34 is evidence rather than a restatement.
Droplets with no biology in them
Douady and Couder dripped magnetised ferrofluid into a dish of silicone oil, and got spiral phyllotaxis with Fibonacci parastichy numbers out of a system containing no cells, no genes and no plant. That is the strongest evidence the pattern is physics — and the clearest warning about what a model can claim.
The counts change with radius
The same head gives 13 and 21 near the centre, 21 and 34 further out, 34 and 55 beyond that, and 55 and 89 at the rim. The transitions are at computable radii, and a photograph captioned with one pair is a statement about one annulus rather than about a flower.
The Fibonacci ladder
Lower the rise on a cylinder and the parastichy pair climbs — 1 and 2, then 2 and 3, then 3 and 5 — each rung the sum of the two before it. Nothing in the arithmetic mentions Fibonacci, the transitions sit at computable rises, and consecutive ones stand in the ratio 1/φ².
A disc is a cylinder
Vogel's seed head makes the rise fall as one over radius squared, so a disc is not one lattice but a family of them. Feed that into the cylinder's ladder and it predicts where a sunflower's spiral counts change — with nothing fitted, and against a counter that never sees either model.
A cone has a rise that falls
A stem holds one parastichy pair for ever and a seed head changes its pair with radius. A cone does both — it is a cylinder whose rise falls as one over the distance from the apex, and the same blind counter that finds one answer up a stem finds four up a cone.
A pattern with a rate
Every lattice on this site is a static object indexed by a parameter, and a plant is not. Put the rise on a clock, place each node where the repulsion from the ones below it is least, and the object that comes out has a history — which is the first thing here that could disagree with the ladder.
Half the golden angle
The forks of the van Iterson tree converge on 137.5078° and sit at none of them. Divide the whole tree by two and the same thing happens at 68.7539° — which is where teasel is, and where a bijugate sunflower counted 42 and 68 has to be.
The lag that is not there
A pattern built out of its own history should hold its old parastichy pair past the point where a fresh lattice would have changed, and the gap should grow as the shoot is hurried. Over a fifteenfold range of rate it does not — every transition lands within a tenth of a rung of where the static ladder puts it.
Transitions a factor of φ² apart
The ladder's rungs are a factor of 1/φ² apart in rise. A disc's rise falls as one over radius squared and a cone's as one over distance, so the same rungs land a factor of φ apart on a seed head and a factor of φ² apart on a cone — measured, on both, by a counter that has never heard of either.
The shape and the law
A cone's transitions are a factor of φ² apart and a disc's a factor of φ, and the temptation is to read the ratio as the shape. It is not. Five surfaces built and counted show that the ratio measures one exponent, and that the exponent is the shape multiplied by the way material arrives.
Why a cone can be counted once
A pineapple is described as 8 and 13 and the description holds. A sunflower is described as 34 and 55 and the description is a statement about one annulus. Both organs have the same ladder in element number — what differs is where an organ puts its elements.
An organ has no single exponent
The previous phase measured that a surface whose circumference grows as a power of arc length puts its transitions a fixed factor apart, and checked it on five surfaces. Every one of them had a single exponent, and no organ does — a fir cone is an ogive, whose exponent runs from 2 at the tip to nearly 0 at the shoulder.
What one exponent reports
Fit a single shape exponent to an organ that has four of them and it returns a real quantity — the harmonic mean of what its individual steps report. Harmonic means sit below arithmetic ones, so the fit understates, systematically, in a known direction, and invisibly.
How much of a cone to measure
Two rings cannot show a varying exponent — not with difficulty, but in principle, because one gap determines one exponent with nothing left to disagree. Four or five can, if each is found to within a per cent. At three per cent this specimen cannot be told from a power law however many of its rings are recorded.
Named alongside it
The objects these essays reach for when they reach for this one.
RiseLadderBranchConeParastichyFibonacciOgiveCylinderDivergence angleParastichy pairSpecimenThe local exponent