Phyllotaxis
the spirals are counted, not admired
Concept

Convergence — where it appears

Whether a computed quantity stops moving as the effort spent on it is increased. It is checked here by raising the grid, the run length or the number of seeds and requiring the answer to stay, since a number that has not converged is a number about the method.

Named by 12 essays across 3 fields — each of them below, with the objects they name alongside it.

The rule, 26 steps in, at a growth of 0.40. The next primordium goes where the repulsion is least — the marked minimum at 216°. Nothing in the rule refers to any particular angle.

The angle is an output

137.5° is not a constant of nature. It is where a rule settles — a rule that places each new element as far as it can from the ones already there, contains no reference to the golden ratio, and reaches the same answer from starting angles a hundred and sixty degrees apart.

emergence · Attractor Where this implementation stops converging. Below about G = 0.18 the settled angle wanders over 113° however long the run. That is the model's limit, not a fact about plants.

Where the model stops

Below a growth parameter of about 0.18 this implementation does not converge — the settled angle wanders over a hundred degrees however long the run. That is the range where the literature says the interesting behaviour lives, and it is worth a figure rather than a quietly chosen axis.

emergence · Modellimit Fibonacci at a rise of 4.8e-3, asked two ways. Choose a divergence at random and a static lattice at this rise gives a consecutive Fibonacci pair 10.8 per cent of the time. Start coarse at a divergence nobody chose, grow the stem down to the same rise, and it is 100 per cent of 16 runs. The geometry is not generous; continuity is.

Continuity from a coarse start

At a fine rise, one divergence in seven gives a Fibonacci pair. Grow a stem from a coarse start at a divergence nobody chose, down to that same rise, and sixteen runs out of sixteen end on 8/13. The earlier work's interpretation was that continuity does the work; this is the measurement it never had.

emergence · Continuity The fork the cost chooses at a daughter ratio of 1: 37.47 and 37.47 degrees. Three ends held fixed, three weights fixed by the radii, and the branch point put where the total cost is least. At the optimal radius the pumping term is exactly half the upkeep term, so a segment's weight is its own cross-section and the three weights here are 1.5874, 1.0000, 1.0000. Minimising directly over the position — a 41×41 grid re-centred and shrunk 220 times, told nothing about any formula — puts the daughters at 37.4673° and 37.4673° from the parent's own forward direction, against the closed form's 37.4673° and 37.4673°.

The angle the cost chooses

Murray's exponent falls out of minimising a cost over the radius of a tube. The same cost minimised over the position of the branch point instead fixes both fork angles, and 281 networks minimised on a grid told nothing about any formula agree with the closed form to under two ten-thousandths of a degree.

branching · Fork angle Which neighbours decide where an element goes. Each line is one exponent: how much each shell of neighbours makes the energy profile vary around the circumference, divided by what the nearest shell contributes. At p = 0.5 the nearest shell leads the next by a factor of 1.1 and a node is placed against the whole neighbourhood at once. At p = 3 it leads by 9.7e+3, and a node is placed against its immediate neighbours — which is what a lattice is.

How far a primordium reaches

The placement rule's repulsion falls as an inverse cube because that is what two magnetised droplets do, and nothing about a plant supplies the exponent. Asking what it controls produced one tidy wrong answer and one measured right one — and the difference between them is the difference between a total and a variation.

emergence · The range of the interaction One planar crown sized for a load on its tips and for the weight of its own wood. The same symmetric crown, 9 generations deep, each branch 2^(−1/2) the length of its parent and turned 30° at every fork, sized so that every branch is bent to one stress, drawn to one trunk width. On the left the load is on the tips and the trunk junction conserves r to the power 1.980; on the right the load is the weight of the wood, found by iterating the radii until they stop moving, and the trunk junction conserves r to the power 0.969. A crown sized for its own weight thins much faster from the trunk, because a branch's weight grows with the square of its radius.

A crown that carries its own wood

Sizing every branch so that equal loads on the tips bend it to one stress gives a crown filling a plane Da Vinci's exponent of two. Move the load onto the wood and the sizing becomes a fixed point, because a branch's load now depends on the radii being solved for. Under the wind on its wood a planar crown still conserves two, but only as a limit its trunk is two tenths short of at fifteen generations. Under its own weight it conserves one — radius rather than area, the stress-similarity law that radius goes as length squared — and a crown carrying leaves and wood reads the leaves' two near its twigs and the wood's one at its trunk, with the handover set by how much of the trunk's load the wood carries.

branching · Murray At 384 azimuths the ratio is 0.62; converged it is 0.82. The comb ratio and the recorded divergence scatter at a rise of 0.005, against how finely the rule samples the circle when it takes its minimum. At 384 azimuths — the grid every flat run in these essays uses, and the grid that earlier work's 0.65 was measured on — the step is 0.94°, which is larger than the 0.25° disturbance the stems carry. The quantisation is white noise, it dilutes both combs, and it does not dilute them equally. The ratio settles at 0.82 from 1152 azimuths up, and the scatter loses 0.19° that belonged to the grid rather than to the stem.

The grid was in the number

The rule places each organ at the least of a profile sampled at a fixed number of azimuths, and every flat run in these essays samples 384 of them — a step of 0.94°, against a disturbance of a quarter of a degree. The quantisation is the larger of the two, it is white, and it moves the discriminator from 0.79 to 0.62. The convergence study this collection had asked for and never done, in the place it turned out to matter.

cylinder · Instrument ceiling Sizing for equal bending and sizing for equal stress cross at one length ratio. Equal stress holds r³ against the sum of a load's arms and gives 3/(1 + ℓ); equal deflection holds r⁴ against the sum of the arms squared and gives 4/(1 + 2ℓ), with ℓ = log₂(1/λ). Setting them equal gives 3(1 + 2ℓ) = 4(1 + ℓ), whose only root is ℓ = 1/2 — the crown that fills a plane, λ = 0.707107 — and there both are exactly two. Below that ratio the stiffness rule reads the lower exponent of the two and above it the higher, so the two criteria size the same crown at one length ratio in the whole family and it is the one Da Vinci's rule names.

A crown sized for how far it bends

Stress is one criterion for sizing a branch and stiffness is another. Holding every branch to the same deflection as a share of its own length sizes r to the fourth against the sum of each load's arm squared, where equal stress sized r cubed against the arm, and the junctions of a deep crown then conserve 4/(1 + 2·log2(1/λ)). A single cantilever under its own weight comes out at radius as length to the three halves — McMahon's elastic similarity, fitted here rather than assumed — against the square that equal stress asks for. And the two criteria agree at exactly one length ratio out of the whole family: λ = 2 to the minus a half, the crown that fills a plane, where both give exactly two.

branching · Murray How many organs a stem needs before it is on a lattice. One row per rise, one mark per starting angle, placed at the organ from which every later divergence stays within a degree and a half of the run's own final value. Where a stem settles at all it does so between 0 and 290 organs in, against the 400 every ablation run here grows before it cuts anything. Not one row needs the length it is given. What changes down the table is the count on the right: how many of the nine starting angles reach a lattice at all, which falls from 7 at the coarse rises to 1 at the finest.

How long a stem takes to settle

Every result here is grown on a stem that has settled onto a lattice, and settling has always been tested for and never timed. Timed, it takes between nothing and two hundred and ninety organs — against the four hundred every ablation run grows before it cuts anything, and the nine hundred the noise runs carry.

cylinder · Settling The settled divergence down the golden branch. Every rise from 0.07 down to 0.00482, plotted against the divergence the rule settles on, with each rung drawn in its own stroke and the branch's limit angle marked. The curve does not slide: it turns three times in four rungs, climbing across one and falling across the next, so a value it takes on one rung it takes again on another. That is what makes a matched pair possible — two rises, different counted pairs, one angle — and it is the whole reason the design exists on this branch. The widest excursions from the limit angle, coarse rung first, are 3.195°, 3.352°, 0.961°, 0.422°.

The angle the ladder returns to

Down the golden branch the settled divergence climbs across one rung and falls across the next, turning three times in four rungs. That is why the same angle is reached at two different rises — and why the Lucas branch, which turns once, almost never offers the same thing.

cylinder · Same angle How often a stem settles, at three samplings of the starting angle. The share of runs that reach a lattice, over the whole table of four falloff exponents by eight rises. The upper three bars are the pooled share at nine, twenty and forty starting angles; the lower three are the share for each group of angles on its own. The pooled figure falls 40.6 to 32.3 to 30.0 per cent, by 8.3 points and then 2.3, so it is converging. The eleven angles added at twenty and the twenty added at forty settle at 25.6 and 27.7 per cent, which differ by less than their own error.

Forty angles, and a limit

Nine starting angles turned out to be a biased sample of the circle, and doubling to twenty said by how much. Doubling again says the estimate is converging — to a smaller correction than one doubling extrapolated to.

emergence · Settling The widest basin, at twenty starting angles and at forty. The starting angles of one cell of the table, drawn as ticks, with the run of consecutive angles that all reach one destination filled. At twenty angles it holds 7 angles and spans 43.75 degrees; at forty it holds 14 and spans 47.5. Halving the spacing doubled the count and left the width where it was, which is what a real basin does and what a run of angles produced by the sampling does not. The other half of the same table goes the other way: 170 of 233 runs are a single angle, against 116 of 151.

A basin that doubled

A run of consecutive starting angles reaching one destination is a basin, and its width is a lower bound. Halving the spacing doubled the angles in the widest one and left its width alone, which is what a real basin does and a sampling artefact does not.

emergence · Settling

Named alongside it

The objects these essays reach for when they reach for this one.

Honest limitsClaim testingDivergence angleRiseAttractorDiscretisationMeasurementRepulsionBasinBranchClosed formFibonacci

All concepts