Depth

Ladders — page 2

A field says what an essay is about. A ladder says what else there is to say about it — the distinct arguments that stand against one idea, from the one that introduces it to the one that assumes all the others.
5/107.9%5 × 1/23/67.9%3 × 1/22/46.1%2 × 1/24/85.5%4 × 1/27/145.2%7 × 1/26/122.8%6 × 1/28/160.1%8 × 1/2share of divergences at this riserise 0.008 · 3600 divergences35.4% share a factor

What "whorled" was hiding

The expansion phase's census put 35% of divergences in a bucket labelled whorled and moved on. Opened, every pair in it is k and 2k — the coarsest rung of the ladder, repeated k times — and reading the census up to jugacy takes the Fibonacci share from 14.7% to 50.1% without describing a single extra plant.

1 rung · wrong
646668701234567fork number down the treedivergence at the fork (°)137.5078/2 = 68.7539°2-jugate · 7 forkslast fork 68.7365°

Half the golden angle

The forks of the van Iterson tree converge on 137.5078° and sit at none of them. Divide the whole tree by two and the same thing happens at 68.7539° — which is where teasel is, and where a bijugate sunflower counted 42 and 68 has to be.

1 rung · lattices
0.2500.5000.75011.25-2.50-2-1.50-1-0.500log₁₀ of the rise between nodes (falling to the right is the plant growing)log₁₀ of the larger parastichy number2/33/55/88/1313/21500 rises, shortest vectors recomputed at eachratio 0.3820 against 1/φ² = 0.3820

The Fibonacci ladder

Lower the rise on a cylinder and the parastichy pair climbs — 1 and 2, then 2 and 3, then 3 and 5 — each rung the sum of the two before it. Nothing in the arithmetic mentions Fibonacci, the transitions sit at computable rises, and consecutive ones stand in the ratio 1/φ².

1 rung · cylinder
divergence 137.508°closest pair 1.60 × mean spacing

A head is a set of points

The nth primordium at n times an angle, and a radius of root n. Two lines of arithmetic produce a sunflower head, which is either remarkable or suspicious depending on how carefully the claim is stated — and stating it carefully is most of the work.

1 rung · lattices
00.500155.5066.507sides of the cellarea, as a multiple of the mean cell areaLewis269 interior cellsslope 0.014 against 0.25

Lewis's law wants disorder

Cell area rises linearly with side count — measured on cucumber epidermis in 1928 and quoted ever since as a property of packed tissue. It holds beautifully on a random point set, with a fitted constant of 1.64 against Lewis's 2. On a phyllotactic head it does not hold at all: the slope is 0.009, and area and side count are almost independent.

1 rung · tissue
491 symbols · 130 segmentsa description, not a mechanism

L-systems describe, they do not explain

Two rewriting rules and a turtle produce something indistinguishable from a plant, and there is no plant in it — no light, no water, no auxin, no mechanics. That is worth demonstrating precisely because the output is so convincing.

1 rung · branching
120° — a third of a turn3 and 6 — whorled137.51°2 and 3 — Fibonaccirise 0.055 in both panelsthe counts decide, not the eye

What a mechanism would have to show

This site says of every model it draws that reproducing a pattern is not explaining it. That is easy to repeat and hard to make precise. Here it is made precise — a list of what an account of phyllotaxis would have to establish, with each item marked according to whether the models on this site establish it.

1 rung · mechanism
0501000.2500.5000.7501growth parameter Gspread of the last 30 steps (°) — 0 means settledfilled dark: convergedthe usable range is stated, not implied

Where the model stops

Below a growth parameter of about 0.18 this implementation does not converge — the settled angle wanders over a hundred degrees however long the run. That is the range where the literature says the interesting behaviour lives, and it is worth a figure rather than a quietly chosen axis.

1 rung · emergence
branch angle 30° · daughter ratio 1.00127 segments · 63 junctions checked

The cube law

A branching network built to move fluid for the least work obeys one relation at every junction — the cube of the parent radius equals the sum of the cubes of the daughters. It is a minimisation result, it is checkable on a real tree, and it is the rare biological rule with a derivation.

1 rung · branching
golden — 6.85× per turnnautilus — 3.2× per turnsame construction, same start2.14× apart in growth

The nautilus question

A golden spiral grows by 6.854 per turn. Measured nautilus sections give about 3.2. That is a factor of 2.14 — not a rounding error, not an artefact of where the centre is assumed to be, and not close.

1 rung · shells
024680.2000.4000.6000.8001counting only outside this fraction of the organ's length or radiustransitions inside the counted partdisc: 2 beyond 50%cone: 1 beyond 50%flare 0.12 · 4000 elements1 against 2 in the outer 50%

Why a cone can be counted once

A pineapple is described as 8 and 13 and the description holds. A sunflower is described as 34 and 55 and the description is a statement about one annulus. Both organs have the same ladder in element number — what differs is where an organ puts its elements.

1 rung · cylinder
00.2500.5000.7501120130140150divergence angle (°)closest pair, as a fraction of the mean spacing (higher is better)137.508°400 points per anglethree criteria, three winners

Packing, measured four ways

The claim is that the golden angle packs best. It is measurable, and the measurement gives three different winners on three criteria — all near 137.5° and none of them it. That does not make the claim wrong; it makes the usual statement of it wrong.

1 rung · tissue
W = 2.40 · D = 0.42W·D = 1.01 — evolute

Raup's three numbers

Nearly every coiled shell that has ever existed is a point in a three-dimensional space — how fast the whorl expands, how far the opening sits from the axis, how far it travels along it. That is a remarkable compression, and the most useful line in the space is the one where the whorls come apart.

1 rung · shells
-0.075-0.050-0.02500.025510152025number of peaks around the ringgrowth rate of that mode8 counted8 predictedDₐ = 0.000008, D_h = 0.00064predicted 8, counted 8

Turing's last problem

Turing's final work was on phyllotaxis and it was unpublished when he died. Its core is a ring of cells and two diffusing substances, and the thing it does is select a number of peaks — which can be predicted from the equations before anything is integrated, and then counted from what the integration produces.

1 rung · mechanism
the first of the four — 300 of its 899 pointsused to buildcountsrecovered137.508°55 · 89137.520°99.502°47 · 7699.500°151.100°31 · 81151.105°77.960°37 · 6077.960°worst error 0.012°counts in, angle outthe recovery never sees the angle

Recovering the angle from the counts

Build a head at a stated divergence angle, forget the angle, and get it back from the spiral counts alone. Four angles, worst error twelve thousandths of a degree — and the only thing that crossed between the two halves was a list of coordinates.

1 rung · lattices
137.5°disorder 3177°8 peaksdisorder 747°8 peaksdisorder 11109°7 peaksdisorder 19151°7 peaksdisorder 2347°7 peaksdisorder 31133°8 peaksthe angle a spiral would need to repeatsix runs, identical equationsspread 129°

A ring cannot make a spiral

The peaks on a Turing ring do not all appear at once — there is a first and a second. But which two lead is decided by the starting disorder, so the angle between them comes out at 177°, then 47°, then 109°, then 151°. A divergence angle is a relationship that repeats, and this one does not.

1 rung · mechanism
501001501020304050radius in the disclarger parastichy number, measured on the disc21/3434/5555/8989/144prediction from the cylinder, counts from the disc15 of 16 bands agree

A disc is a cylinder

Vogel's seed head makes the rise fall as one over radius squared, so a disc is not one lattice but a family of them. Feed that into the cylinder's ladder and it predicts where a sunflower's spiral counts change — with nothing fitted, and against a counter that never sees either model.

1 rung · cylinder
1/2 at the bottom, 5/8 at the top167 nodes · rise 0.2 → 0.004542 nodes per rung

A pattern with a rate

Every lattice on this site is a static object indexed by a parameter, and a plant is not. Put the rise on a clock, place each node where the repulsion from the ones below it is least, and the object that comes out has a history — which is the first thing here that could disagree with the ladder.

1 rung · emergence
1231234rise exponent p, where the rise falls as z⁻ᵖratio between consecutive transitions along the axis5 surfaces, 16 measured transitionsworst disagreement 0.8%

The shape and the law

A cone's transitions are a factor of φ² apart and a disc's a factor of φ, and the temptation is to read the ratio as the shape. It is not. Five surfaces built and counted show that the ratio measures one exponent, and that the exponent is the shape multiplied by the way material arrives.

1 rung · cylinder
5 sides13420%6 sides45869%7 sides7311%665 bounded cellsmean 5.908 sidessix is forced, not chosen

Why the average cell has six sides

Not because hexagons are efficient. Because Euler's formula leaves a tiling no choice — count the edges two ways and the mean comes out at six, whatever the cells would prefer. The efficiency argument is a different claim about a different thing.

1 rung · tissue

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