Phyllotaxis
the spirals are counted, not admired
Concept

Initial condition — where it appears

The stretch of pattern a run is started from, which puts it on one branch rather than another. It is a parameter of every result here, and treating it as one is what turns which pattern appears into a question with an answer.

Named by 11 essays across 3 fields — each of them below, with the objects they name alongside it.

The angle between the first two peaks, for six starting disorders. The same equations, the same ring, six different starting perturbations: 177°, 47°, 109°, 151°, 47°, 133°. A spiral needs the angle between successive elements to be the same one each time, and a stationary ring does not supply that.

A ring cannot make a spiral

The peaks on a Turing ring do not all appear at once — there is a first and a second. But which two lead is decided by the starting disorder, so the angle between them comes out at 177°, then 47°, then 109°, then 151°. A divergence angle is a relationship that repeats, and this one does not.

mechanism · Ring limit One seed, two rates, two ladders. Both stems begin as forty nodes of Lucas lattice at a rise of 0.12. At 65 nodes per rung the divergence stays at 99.5° and the counts walk 1/3 → 3/4 → 4/7 → 7/11. At 131 it leaves for 137.7° and walks 1/3 → 2/3 → 3/5 → 5/8 → 8/13 instead.

The rate decides the branch

Forty nodes of Lucas lattice, carried down to the same fine rise twice. Hurried, the pattern holds the Lucas ladder through three more forks at 99.5°. Given room, it abandons it at the first fork it reaches and walks to 8/13 at 137.5°. Same seed, same rise, two ladders — with a threshold between them at about ninety nodes per rung.

emergence · Branch choice Fibonacci at a rise of 4.8e-3, asked two ways. Choose a divergence at random and a static lattice at this rise gives a consecutive Fibonacci pair 10.8 per cent of the time. Start coarse at a divergence nobody chose, grow the stem down to the same rise, and it is 100 per cent of 16 runs. The geometry is not generous; continuity is.

Continuity from a coarse start

At a fine rise, one divergence in seven gives a Fibonacci pair. Grow a stem from a coarse start at a divergence nobody chose, down to that same rise, and sixteen runs out of sixteen end on 8/13. The earlier work's interpretation was that continuity does the work; this is the measurement it never had.

emergence · Continuity What the model settles on, against how fast the meristem grows. A broad golden branch, a transition, and then the two-whorl regime at exactly half a turn. 16 of 40 converged settings land within 4° of the golden angle; 17 land more than 20° away.

Two-ranked, by two different routes

The rule produces a two-ranked stem at a coarse rise, where 180° is the only thing available, and that has been in the bifurcation diagram since the beginning. It also produces one at a fine rise, at a rise whose own answer is the golden angle, if a single organ is removed. The diagram cannot show the second, and the reason it cannot is how it is drawn.

mechanism · Attractor One rule, one rise, two branches that stay where they were put. The top 70 organs of two stems grown by the same placement rule at the same rise of 0.013, differing only in the stretch of ideal lattice each was started from. The left one was seeded at the golden angle and settles at 136.781° with the pair 5/8; the right one was seeded on the Lucas lattice and settles at 99.785° with 4/7. Neither drifts towards the other: 0.73° and 0.28° from where each was seeded, over four hundred organs. That is what makes an intervention on the right-hand stem a measurement about a different lattice rather than about a different rule — and 4 and 7 are not Fibonacci numbers, which is the property the experiment needs.

A stem on the other branch

Every stem an organ had been cut from carried Fibonacci counts, which is why two rival explanations of the block a wrecked stem settles into had never disagreed. A stem seeded on the Lucas lattice carries four and seven at the same rise, under the same rule. Cut, it settles on seven — the larger number, and not a Fibonacci one.

cylinder · Branch choice The wrecked stem is the lattice it was cut from, wound the other way. The divergence of each organ placed after two were removed from a settled stem at a rise of 0.032, over the 220 organs following the cut. The upper line is the divergence the undisturbed stem holds, 139.6875°; the lower is 220.3125°, which is 360° minus it and therefore the same lattice with the opposite handedness. The sequence is thrown by the cut, wanders for a few dozen organs, and settles on the second line to four decimal places — 220.3125° against 220.3125° — where it stays. A counter shown the positions afterwards returns 3 and 5, the pair the stem was cut from. Nothing about the pattern has been lost; its chirality has been reversed, which at this rung a single removal cannot do.

The stem that changed hands

A stem that never recovers from an ablation is supposed to end up somewhere worse than it started — a repeating block of angles, a pattern with the wrong counts in it. At the coarsest arrangement it ends up somewhere that is not worse at all: at 220.3125°, which is 360° minus the divergence it was cut from. The lattice is intact and its handedness is reversed.

emergence · Attractor How many organs a stem needs before it is on a lattice. One row per rise, one mark per starting angle, placed at the organ from which every later divergence stays within a degree and a half of the run's own final value. Where a stem settles at all it does so between 0 and 290 organs in, against the 400 every ablation run here grows before it cuts anything. Not one row needs the length it is given. What changes down the table is the count on the right: how many of the nine starting angles reach a lattice at all, which falls from 7 at the coarse rises to 1 at the finest.

How long a stem takes to settle

Every result here is grown on a stem that has settled onto a lattice, and settling has always been tested for and never timed. Timed, it takes between nothing and two hundred and ninety organs — against the four hundred every ablation run grows before it cuts anything, and the nine hundred the noise runs carry.

cylinder · Settling The same table, grown 2.7 times as long. Every rise and every starting angle, grown to 1200 organs and then to 3200. The two middle columns are how many starting angles reached a lattice at each length, and they are the same column: of the 72 pairs of runs, 72 are identical organ for organ and 0 settle at the longer length after failing at the shorter one. Tripling the budget buys nothing anywhere. What the fine rises are short of is not run length: the share of starting angles that reach a lattice at all falls from 7 of 9 to 1, so the arrangements a stem could fall into have mostly stopped existing.

A wall and not a budget

Below a rise of about 0.005 this collection's stems stop settling onto a lattice, and the limit has been written up four times without anybody asking which kind of limit it is. Grown three times as long, the table is identical row for row: not one stem that failed to settle succeeds. The floor is a wall.

cylinder · Settling Every divergence a stem settles on, by falloff exponent. One row per exponent, one mark per distinct settled divergence anywhere in that column, read to a tenth of a degree. This is the sharpest instrument in the sweep and the cheapest: it is already in every run, it does not average, and it is read on the same azimuth grid at every exponent. The steeper falloffs reach 42.3°, 47.9°, 148.1° — values that appear nowhere in the two shallower columns — while the traffic the other way is empty. So the exponent changes the map of where a stem can end up while leaving alone the share of starting angles that end up anywhere, which is a result a measurement of "did it settle" was never going to find.

Destinations only a steep rule reaches

A settled stem's divergence is the sharpest instrument in the sweep and the cheapest: it is already in every run, it does not average, and it is read on one grid at every exponent. Exponents 4 and 5 reach 42.3°, 47.9° and 148.1°, and the two shallower ones reach none of them.

emergence · Settling Every destination in the settling table, counted. One row per destination, placed across at the divergence the stems settled on and labelled with the parastichy pair a counter returns at the top of the run. 117 settled runs land on 15 destinations when two values within half a degree are called one. five of them sit on the golden or the Lucas sequence and ten do not, and the ones that do not are reached at every falloff exponent including the shallowest. Nothing here refuses to count: the settling test does not admit arrangements this collection would decline to call lattices.

A counter on the settling table

The settling table has reported an angle for every run that reaches a lattice, and nobody has ever counted one. A hundred and seventeen settled runs, regrown and counted: every one of them has a parastichy pair, and sixty-five of them land on sequences the ladder does not carry.

emergence · Settling What the 3 steep-only destinations count as. Each block is a divergence that only the two steeper falloff exponents settle a stem on, with the pair a counter returns and the sequence that pair belongs to. All three count. None is a rung of either ladder. Two of them are the only members of their sequence anywhere in the table; the third, at 148.1 degrees, is the coarsest rung of a sequence that supplies another destination at every exponent — so it is not a new kind of arrangement but a coarser one of a kind the table already reached.

What a steep rule counts as

Exponents four and five settle stems on 42.3°, 47.9° and 148.1°, and the two shallower ones reach none of them. All three count: 8/9, 8/15 and 2/5. None is a rung of either ladder, and one of them is the coarsest rung of a sequence the table already had.

emergence · Settling

Named alongside it

The objects these essays reach for when they reach for this one.

AttractorHonest limitsMeasurementRiseDivergence angleBasinThe placement ruleLadderCounting blindFibonacciLatticeNegative result

All concepts