What a plant might be doing

Removing a neighbour costs least

Take away an organ that is a direct chain-neighbour of the growing tip and the next organ moves by under thirty-one degrees. Take away anything else inside the front and it moves by at least sixty-three. Thirty cuts, two groups, a factor of two between them and nothing in the gap.

Worth reading first: The organ that was taken away · A head is a set of points · Counting the spirals.

Every ablation table in this collection carries a number nobody has read: the displacement of the first organ placed after the cut. It is how the front is measured — an organ that moves is inside the front and one that does not is outside — and beyond that threshold nothing has been done with it.

Split the thirty wrecked cuts by which organ was taken away, and the number separates into two groups with a gap between them.

What a removal costs the next organ. One mark per wrecked cut in the census: how far the first organ placed after the removal ended up from where the control put it. The rows split by which organ was taken. Removing a direct chain-neighbour of the growing tip — an organ at a multiple of one of the two counted numbers — moves the next organ by between 8.9 and 30.7 degrees. Removing anything else inside the front moves it by between 62.8 and 167.6. Nothing lands between the two groups and the ratio across the gap is 2.05, so the line is a gap rather than a threshold. Taking away a neighbour is the cheap removal, which is the opposite of what the words suggest.
Fig. 1 Every wrecked cut, placed by how far the next organ moved, with the rows split by whose neighbour was removed.

Remove a direct chain-neighbour of the growing tip — an organ at a multiple of one of the two counted numbers — and the next organ moves by 8.9° to 30.7°. Remove anything else inside the front and it moves by 62.8° to 167.6°. Nine rows against twenty-one, a factor of two across the gap, and nothing lands in it.

That is the opposite of what the words suggest, which is the reason to write it down. It is also a fact about the placement rule rather than about plants, and this collection is careful about the difference: what follows is an account of what a stated rule does when an organ is taken out of the arrangement it is placing against, and any claim about a meristem would need the rule to be the mechanism, which nothing here establishes.

Why it sounds backwards

The placement rule puts each new organ where the sum of inverse powers of distance to the organs already there is least. The nearest organs contribute the largest terms. So removing one of the nearest organs removes one of the largest terms in the sum, and the obvious expectation is that this is the most disruptive thing a removal can do.

The rule, 26 steps in, at a growth of 0.40. The next primordium goes where the repulsion is least — the marked minimum at 216°. Nothing in the rule refers to any particular angle.
Fig. 2 The rule the expectation comes from: a sum of repulsions, with the nearest contributors weighted hardest.

The measurement says the opposite. Removing a chain-neighbour is the cheap removal, by a factor of two, with no overlap between the groups at all.

The next organ moves for the last 8, and for no others. One row per organ removed, counted back from the tip of a stem at a rise of 0.013 whose counted pair is 5 and 8. Removing any of the last 8 moves the next organ by 4.9° to 164.1°; removing an older one moves it by at most 0.70°, which is under the azimuth grid. The boundary is at 8, and 8 is the larger parastichy number — so the experiment counts the spirals without measuring an angle.
Fig. 3 The same quantity at every offset on one lattice, which is the table the two groups are cut out of.

The account

It is a statement about where the removed organ sits in azimuth rather than about how large its term was.

The new organ goes into the gap the existing organs leave. On a settled lattice that gap is bounded on two sides, and the organs bounding it are exactly the two contact families’ nearest members — at lags p and q, on opposite sides of the new organ’s slot. That opposition is what makes them the contact pair.

A cell's neighbours are its spiral families. Left: part of a 700-point golden, 137.508° head, with every contact between two cells drawn and coloured by the difference between the two nodes' placement indices. Right: the share each difference takes, across all 1459 contacts between interior cells. They are the parastichy numbers — 34, 55, 21, 89, 13, 8 — and the pair a person would count is 34 and 55. The six sides Euler forces are shared out among four of them, 5.64 edges per cell.
Fig. 4 The two organs the new one is placed between, which are the nearest members of the two families.

Now remove one of them. The slot’s two walls become one wall and a much more distant one, so the minimum of the sum slides toward the missing wall — but only until the next member of the same family starts to bound it, which on a lattice is one step further along the same chain and in nearly the same direction. The gap widens; it does not open.

Tracing one family: 10 chains. Every node is joined to the node one repeated displacement away, and the chains that result are drawn separately. There are 10 of them, which is the parastichy number of this family. No index of arrival was used anywhere, which is what lets the same count be made on a pattern where 2 primordia appear at once.
Fig. 5 A chain through the arrangement, whose members bound the slot from one side one after another.

Remove an organ that is on neither chain, and the geometry is different. Such an organ sits between the two walls in azimuth — not bounding the slot but sitting across it, several rows down. Removing it opens the interior of the region the new organ is choosing within, and the minimum can move a long way before anything stops it.

Two answers 138° apart, and one organ holding the second one up. The repulsion the rule minimises, around the circumference of a stem at a rise of 0.008, at the height the next organ will sit at. It has two low points 138.3° apart: the slot the next organ takes, and the slot the organ after it will take. The runner-up is 13.6% higher. The organ 13 places back carries 14.6% of the energy at the winning slot and twelve places back carries 16.3% at the runner-up — and that is more than the gap, so taking that organ away makes the runner-up win and the next organ appears a whole divergence away. Neither guard is a contact of the organ being placed; twelve is one place inside the larger number of the pair this stem is climbing towards.
Fig. 6 The slot and its surroundings, where the difference between bounding a gap and sitting inside one is visible.

So the rule of thumb is: a chain-neighbour is a wall and everything else is furniture. Taking away a wall moves the answer to the next wall along, which is close. Taking away furniture lets the answer roam.

The measurement, stated exactly

The number is the azimuth of the first organ placed after the cut, minus the azimuth of the same-numbered organ in a control that shares the stem’s entire history, folded to within half a turn. It is not an average over anything and it has no window in it, which is worth saying because the same runs’ later organs are useless for this kind of question: a wrecked stem is permanently displaced everywhere above the hole, so any quantity read further up is measuring a phase difference rather than a response.

The first organ is the exception. It is placed by the rule against a configuration that differs from the control’s in exactly one organ, so its displacement is a clean response to a single change with nothing accumulated in it.

Take away the organ eight places back, and the next one goes into the hole. The last 30 organs of a stem at a rise of 0.005, unrolled. The open circle is the organ removed — eight places before the tip. The ring at the top is where the rule puts the next organ with every organ present; the filled mark beside it is where the rule puts it with that one missing. The two are 16.4° apart, against a local spacing of 25°, and the vacancy itself is 22.7° from the undisturbed answer. Nothing else differs between the two runs: same rise, same history, same rule.
Fig. 7 The one organ the measurement is about, and the two configurations it was placed against.

That is also why the result is available at all. Every other quantity in this thread is read off a settled stem hundreds of organs later, where the cut’s signature has been overwritten by the new phase. This one is read immediately, and immediacy is what makes it about the geometry of the hole.

What a cut moves, organ by organA stem counted at 5 and 8 spirals with the organ five places back from the tip removed, compared against a control that shares its history to the last digit. Each mark is one organ placed after the cut and how far its azimuth ended up from where the control put the same organ. The quantity folds at half a turn, so 152 degrees is near the largest displacement there is; and it does not decay with height, which is what a stem that never repairs means. There is therefore no organ that the cut disturbed most in any useful sense, and a reading that needs one has nowhere to stand.-200-100010020010203040organs above the removed oneazimuth moved (°)the removed organ was 5 places back5/8 at 0.013 · offset 5 · survivor 5generated from a stated rule, not drawn to look right
Fig. 8 What happens after that first organ, which is why nothing later can be read as a response.

The gap, and why it is a gap rather than a line

Nine measurements between 8.9° and 30.7°; twenty-one between 62.8° and 167.6°. The ratio across the empty region is 2.05.

That matters because a threshold chosen to make a claim true is how a claim gets made true. Nothing here is fitted: any cut-off between about thirty-five and sixty degrees gives the same two groups, and the tolerance stated in the machinery — forty-five degrees — is the middle of that range rather than the edge of it.

What a removal costs the next organ. One mark per wrecked cut in the census: how far the first organ placed after the removal ended up from where the control put it. The rows split by which organ was taken. Removing a direct chain-neighbour of the growing tip — an organ at a multiple of one of the two counted numbers — moves the next organ by between 8.9 and 30.7 degrees. Removing anything else inside the front moves it by between 62.8 and 167.6. Nothing lands between the two groups and the ratio across the gap is 2.05, so the line is a gap rather than a threshold. Taking away a neighbour is the cheap removal, which is the opposite of what the words suggest.
Fig. 9 The two groups with the empty region between them drawn, which is what makes the split a measurement rather than a choice.

It also matters that the grouping is decided by arithmetic on the offset — the same arithmetic the lost-member reading uses to decide which rows it can be asked on — and not by the displacement. Whether a removed organ is a chain-neighbour is settled by whether its offset divides by five or by eight, with no reference to how far anything moved. So the two variables are independent, and their agreement across thirty rows is a result rather than a definition.

Every wrecked offset, and whose neighbour was removed. Each row is a stem that never repaired, with the family of the organ that was taken and the family that survived. An organ five places back on a stem counted at 5 and 8 spirals lies on the tip's five-chain, so the question can be asked there; an organ four places back lies on neither chain and it cannot. Of 30 wrecked offsets in the census, 9 remove a member of exactly one family and 21 remove a member of neither. On every one of the 9 the family that lost a member is the family left standing, which is the opposite of what the reading predicted.
Fig. 10 The arithmetic that does the grouping: which chain, if either, the removed organ lies on.

What it predicts

Three things, and the first two are already visible in tables here.

That the front’s edge should be ragged rather than sharp. The front is the run of offsets at which a removal is felt at all, and it has been reported as a step: large out to the larger counted number and nothing past it. If chain-neighbours are cheap, then the offsets equal to p, q and their multiples should sit lower than their neighbours inside that step — a comb of notches rather than a plateau.

The next organ moves for the last 13, and for no others. One row per organ removed, counted back from the tip of a stem at a rise of 0.005 whose counted pair is 8 and 13. Removing any of the last 13 moves the next organ by 2.6° to 167.6°; removing an older one moves it by at most 0.47°, which is under the azimuth grid. The boundary is at 13, and 13 is the larger parastichy number — so the experiment counts the spirals without measuring an angle.
Fig. 11 The step, at a finer rise, with the notches the account predicts.

That the effect should not depend on the rise except through the pair. The account is about which organs bound the slot, and that is fixed by the counted pair — which is also why a band that holds the pair and moves the rise would be expected to leave it alone. Across the census the two groups hold at every rise from 0.032 to 0.005, which spans a factor of six and four different pairs.

The band that never heals is what two fixed edges leave over. Each row is one rung. A stem is grown to 400 organs, one organ is removed, and the stem is followed for 300 more. A filled mark is an offset the stem recovers from, with the number of organs it took printed above it; an open ring is an offset it never recovers from. At the 3/5 rung, a front of 5, every offset heals. At the 5/8 rung, a front of 8, the offsets 4, 5 never heal. At the 8/13 rung, a front of 13, the offsets 4, 6, 7, 8, 9 never heal. What is the same at every rung is the two edges: the first three offsets heal, and so do the last few of the front. What is left in the middle is the band, and it widens because the front does.
Fig. 12 Three rises with three pairs, where the same split holds in each.

That a two-organ cut removing both walls should be much more expensive than either alone. That is testable with machinery already here and it has not been done: the multi-organ work measures whether a stem repairs, not how far the next organ moves.

Move the second organ far enough back and the experiment is the old one. The displacement of the next organ when two organs are removed — one two places back and one a further gap behind it — against that gap, at a rise of 0.032 where the pattern is 3/5. The dashed line is what removing the single organ two places back does on its own, computed by the earlier one-organ intervention and not by this one. Inside the front the two vacancies interact and the answer swings over 77°; from the gap that puts the second organ 2 places behind the front onwards it settles onto the single cut's 79.0°, within 0.4°. That limit is what makes the second parameter a control rather than a confound.
Fig. 13 The two-organ machinery the prediction would use, which measures a different quantity today.

What it does not explain

It does not explain which family survives. That is the question this thread has been chasing, and the cheap-removal result is about the first organ rather than about the eventual arrangement.

The two are not obviously connected, and the measurement says they are not connected simply. Among the nine cheap removals the survivor is the removed organ’s own family every time — that is the lost-member reading — and among the twenty-one expensive ones the survivor is whatever the offset rule says it is, with the same five exceptions as ever.

Which lag survives, at every lattice and every offset. A row for each of twelve lattices and a column for each offset an organ is removed at, with the lag whose hop survived written in the cell. 30 cells never repair. The lag left standing is one of the two contact families at 29 of them, and at 17 it is the second shortest step on the lattice rather than the shortest, which is what refuses the reading that a rule holds its nearest neighbours. The ringed cells are the five the offset rule gets wrong. Two lattices carry no cells at all because no single removal wrecks them.
Fig. 14 The survivors across the census, which the cheap-and-expensive split does not predict.

So a large first displacement and a small one both lead to a wrecked stem with a rigid hop in it, and how far the first organ moved does not say which hop. If anything that is the more interesting half: the amount of damage and the kind of damage are independent here. It is also a warning against the tempting shortcut of treating displacement as a proxy for severity — the family a stem keeps is not ordered by it, and neither is whether the stem repairs at all, since offsets outside the front move nothing and heal while offsets deep inside it move everything and heal too.

One wrecked stem, lag by lag — golden, rise 0.013, organ 4 back. How much each lattice hop moves from organ to organ in a stem that never repaired after a single removal, over its last 120 organs. The lag-one hop is the divergence itself and it swings by 65 degrees. The lag-5 hop swings by 0.11 degrees and sits 0.09 degrees from where the undisturbed stem put it, so that one family of the original lattice is still standing organ by organ. Its multiples inherit the same steadiness and nothing else comes within a factor of twenty. The block of angles this stem repeats has a period of 5, which is the surviving lag and not a coincidence.
Fig. 15 The kind of damage, read out of a wrecked stem, which the amount does not predict.

The count of nine, and what it rests on

Nine cheap removals is not many, and it is worth saying which nine so the reader can see they are not one lattice’s worth.

They are spread across six lattices and both branches: 5/8 stems at three different rises, an 8/13 stem, two 4/7 stems and a 7/11 stem. Five of them remove a member of the smaller family and four the larger. No lattice contributes more than two.

Which side of the tip the removed organ was on. Each row is one wrecked cut. The centre line is the azimuth the next organ would have taken; the two open marks are where the two contact families leave it, which are always on opposite sides because that is what makes them the two nearest neighbours of a lattice point. The filled mark is the organ that was removed. Reading the lost-member rule as a question about which side rather than about which chain makes it answerable on all 30 cuts instead of 9, reproduces the published reading on every one of those 9, and is right on 22 of 30 overall.
Fig. 16 The census rows laid out by where the removed organ sat, where the cheap ones are the marks sitting on a family’s own azimuth.

That spread matters because the alternative explanation for a clean split is that one lattice happens to behave differently from the rest and its rows have been gathered into a group. Six lattices, two branches and four counted pairs is not that.

What nine rows cannot support is any statement about the shape of the cheap group. Whether removing a member of the smaller family costs more or less than removing one of the larger, whether the cost grows with the offset, whether it depends on the rise — all of those are questions about the internal structure of nine numbers spanning 8.9° to 30.7°, and nine numbers will answer them however they are asked.

What a removal costs the next organ. One mark per wrecked cut in the census: how far the first organ placed after the removal ended up from where the control put it. The rows split by which organ was taken. Removing a direct chain-neighbour of the growing tip — an organ at a multiple of one of the two counted numbers — moves the next organ by between 8.9 and 30.7 degrees. Removing anything else inside the front moves it by between 62.8 and 167.6. Nothing lands between the two groups and the ratio across the gap is 2.05, so the line is a gap rather than a threshold. Taking away a neighbour is the cheap removal, which is the opposite of what the words suggest.
Fig. 17 The nine, drawn beside the twenty-one, at the resolution the claim is made at and no finer.

What it does say about the mechanism

Something narrow and worth having: the rule’s response to a removal is geometric rather than energetic.

An energetic account would predict the response from the size of the term removed, which is largest for the nearest organ. A geometric account predicts it from where the removed organ sat relative to the gap being filled. The two accounts differ in sign here, and the measurement picks the geometric one.

The neighbourhood of 21/55, and where its background was taken from. μ₂ across nine tenths of a degree either side of 21/55, on a head of 825 organs — 15 in each of its 55 rows. The deep notch at the centre is the dip. The shaded columns are the offsets this site now takes a background from: those at least 0.03° from every rational with a denominator of 60 or less, of which there are 54 here. The marked sample at 0.2° is the one the earlier work used, and it lands 0.007° from 13/34 — inside that rational's own dip. It reads 0.115 against a floor of 0.148, so measured that way the dip is an inversion. The clear offsets give 0.339.
Fig. 18 The terms an energetic account would weight by, whose ordering does not match the outcome.

That is a mild result and it is stated mildly. It does not follow that energy is irrelevant — the rule is an energy — only that the response to a perturbation of it is dominated by the shape of the constraint set rather than by the magnitude of the perturbation. In a system where the minimum sits in a narrow valley, removing a wall moves the minimum; removing a distant contributor deepens the valley without moving it much. Here it is nearly the reverse, and it is the reverse because the “distant” contributors are the ones sitting inside the region the minimum lives in.

Three cut-offs at the same nominal width of 3 spacings. The weight the interaction is multiplied by, against distance. They halve at 2.08 (exponential), 2.50 (gaussian), 3.00 (hard) spacings — so a rule described as "cut off at 3 spacings" is three different rules until the falloff is named. Every later figure is read in half-weight radii for that reason.
Fig. 19 The neighbourhood shapes this reasoning is stated over, drawn as the rule sees them.

Where the number was hiding

This collection has had the thirty numbers since the census was built, and every one of them has been printed in a figure. The reason nobody read them this way is worth a paragraph, because it is the ordinary reason.

The first displacement was introduced as an instrument: a way of deciding whether an organ is inside the front. For that purpose all it needs is a threshold, and the threshold is generous — two and a half degrees, chosen because inside the front the displacements run from a few degrees to a hundred and sixty and outside it they are under half a degree. Once a quantity is being used as a detector, its value stops being interesting; what matters is which side of the line it falls on.

The next organ moves for the last 5, and for no others. One row per organ removed, counted back from the tip of a stem at a rise of 0.032 whose counted pair is 3 and 5. Removing any of the last 5 moves the next organ by 8.7° to 149.1°; removing an older one moves it by at most 1.41°, which is under the azimuth grid. The boundary is at 5, and 5 is the larger parastichy number — so the experiment counts the spirals without measuring an angle.
Fig. 20 The quantity as a detector: a step, read for where it falls to zero.

So the number was drawn thirty times, read as a boolean thirty times, and its range — from 8.9° to 167.6°, a factor of nineteen — was never the subject of a sentence. The general form is that a quantity used to make a decision stops being read as a measurement, and the fix is not vigilance but occasionally asking what else a detector’s own output would say if it were a reading.

Both vary; only one of them varies enough to find. Each organ's step exponents, divided by its own mean so the two are comparable. The ogive's run over 15 per cent of their mean across 5 rings. The convex head's run over 1.15 per cent across 5 — inside the band a 3 per cent error on each ring position leaves, so no ruler separates it from a flat disc.
Fig. 21 The habit this belongs to, which is the same one that turned a rise into a fraction of a rung.

What would refute it

A lattice where a chain-neighbour removal is expensive. The account predicts there is none, because the geometry it rests on — two families bounding a slot on opposite sides — is what a contact pair is, on every lattice.

Which offsets give short hops, at a rise of 0.032. The two lowest points are at 3 and 5, and those are the parastichy numbers. Offset 1 is high because a hop of one node is at least the rise, which is what makes a stem easier to count than a disc.
Fig. 22 The ranking that defines a contact pair, whose top two entries the account depends on.

The place to look is the coarse end, where the counted numbers are small and the distinction between “on a chain” and “not on a chain” nearly disappears. On a 2/3 stem, offsets one, two, three, four and six are all multiples of a counted number, so almost every removal is a chain removal and the two groups have nothing to separate. Whether the split survives there is a question about a rung this census does not reach.

Which rises are a lattice, from 0.04 to 0.13. How much the divergence wanders over the last sixty organs, at each rise up the coarse ladder. 13 of the 19 settle, scattering between 0.0000 and 0.3356 degrees. six do not: from 0.09 to 0.115 the divergence sticks on exactly 135.0000 degrees, which is three eighths of a turn, and wobbles about it by 0.79 to 1.60 degrees. A counter shown either kind returns the same pair, so the counts cannot tell them apart. The line is the threshold a rise has to pass before a cut is made on it, and it sits in the gap rather than among the measurements.
Fig. 23 The coarse end where the distinction thins out, and where the census has no rows.

The other place is a whorled stem, where organs arrive several at a time and the notion of a chain through the tip is different. A whorled arrangement’s response to a cut has been measured for whether it repairs and not for how far the next organ moved, so the number is one column away.

three stems: 1, 2, 3 primordia at a time. 1-jugate at 137.51° counts 2 and 3; 2-jugate at 68.75° counts 2 and 4; 3-jugate at 45.84° counts 3 and 6. Every count shares the factor k, and dividing it out leaves an ordinary lattice: what a k-jugate stem is, exactly, is k copies of an ordinary one wrapped k times round.
Fig. 24 The whorled stems where the account would need restating, since a chain through the tip means something else there.

What is left

The notches. If chain-neighbour removals are cheap, the front’s displacement profile has structure in it that every table here has drawn as a step, and reading that structure is arithmetic on data already computed — the offset table exists at three rises and has been read only for where it falls to zero.

On a rung the response is a run of offsets; near a transition it has a hole. One row per rise, from 0.032 at the top to 0.005 at the bottom, and one column per offset: the organ one place back at the left, 14 places back at the right. A cell is filled where removing that organ moves the next organ by more than 2.5°, and empty where it does not. On a rung the filled cells are a run from one to the larger parastichy number — 5, 8, 13 at the pairs shown on the left. Every rise shown here is in the middle of its rung, so every row is a run and nothing past it is felt — what happens between the rungs is a separate matter.
Fig. 25 Those tables, drawn at three rises, read here for their far edge and not for their shape.

And the two-organ test, which is the sharpest thing the account predicts and the only one that would need new runs. Remove both walls and the new organ should move much further than either removal alone; remove two pieces of furniture and it should move about as far as one. That is a two-by-two comparison with an interaction in it, which is more than any single measurement here has asked for.

More organs removed, more stems that never come back. The share of arrangements at the 5/8 rung that never return to the divergence they were cut from, against how many organs the cut removed. One organ wrecks 2 of 8 arrangements and five wreck 63 of 64. The number of arrangements differs from bar to bar because a cut of five organs has more ways of being placed than a cut of one, and it is printed on each bar for that reason. What the dose decides is whether a stem falls off its lattice; where it lands when it does is decided by something else.
Fig. 26 The multi-organ machinery the test would use, measuring today whether a stem repairs rather than how far its next organ went.

Shares its objects with

Essays that name at least two of the same things, and that neither author linked.

Named objects

A flat tag is an object no other essay names yet.

AblationContact networkControlLargest gapLattice offsetMeasurementMechanismNearest neighbourNeighbourhoodParastichyParastichy pairThe placement rulePredictionRepulsion