Concept

Nearest neighbour — where it appears

The organ closest to a given one on the surface, which on a lattice is a member of one of its parastichy families. Distances to neighbours are how a counter recovers a pair without being told anything about the angle.

Named by 39 essays across 7 fields — each of them below, with the objects they name alongside it.

The two spiral families a counter finds between 0.68 and 0.92 of the radius. 34 spirals one way and 55 the other, found from the point positions alone — the counter is never told the divergence angle.

Counting the spirals

Almost every claim about phyllotaxis is a claim about how many spirals run through a pattern, and the count is almost never done. It can be done from the points alone, by a count that is never told what angle built them — and then a count of 34 is evidence rather than a restatement.

lattices · Counting
Closest pair across 120–155° at 400 organs, read both ways. On the interior's scale the golden angle reads 0.9027 and ranks 1st of 72, against 0.9026 for the best grid angle at 137.5°, with the window running from 0.0668 to 0.9026; counting the rim's cells the golden angle reads 0.7076 and ranks 2nd of 72, against 0.7129 for the best grid angle at 137.5°, with the window running from 0.0331 to 0.7129. The dashed upright is the golden angle, which a grid of decimal degrees never lands on and which is therefore read separately.

Packing, measured four ways

The claim is that the golden angle packs best, and it is measurable. Read on the interior of a head, the two criteria about distance put the golden angle first among the angles near it and the two about cells are won by rational angles — which makes the claim half right, and makes the right half a statement about a class of angles. An earlier reading of the same four criteria, divided by the cells at the head's edge, said the opposite.

tissue · Packing
The site's hero shell comes free at a translation of 1.1066, and a slower one at 10.49. Setting the located boundary to zero and solving leaves T_free = √W/(W−1), drawn here across the whole expansion range on logarithmic axes. Above the curve the shell is free at every distance from its axis and the contact region has left the plane rather than merely shrunk in it; a slowly expanding shell at an expansion of 1.1 needs 10.4881 turns of translation to buy that and one expanding twentyfold needs 0.2354. The tower is what buys a shell the right to coil close to its own axis, and the faster it expands the less tower it takes.

What a spire buys

Translation along the coiling axis enters the contact boundary as its square, so a shell that has only just begun to walk along its axis has not moved the boundary at all. It is also strictly one-way, and it has a threshold above which no distance from the axis whatever puts the whorls in touch.

shells · Morphospace
Tracing one family: 4 chains. Every node is joined to the node one repeated displacement away, and the chains that result are drawn separately. There are 4 of them, which is the parastichy number of this family. No index of arrival was used anywhere, which is what lets the same count be made on a pattern where 2 primordia appear at once.

Counting without an index

A person counting spirals on a cone puts a finger on one scale, follows a family round, and counts how many distinct chains there are. That needs no order of arrival — and building it turns out to be strictly more general than the counter that reads the order of arrival, and to find a bug in the counting of a bijugate stem that nothing had caught.

lattices · Chain counting
A lattice with an error inherited from the two contact neighbours. The autocorrelation of 759 divergence angles from a kinematic lattice at a divergence of 137.8261° and a rise of 0.005, with 0.5° of scatter on each azimuth. There is no placement rule anywhere in it: node i is put at exactly i times the divergence and then displaced. The only thing that differs between this figure and the control is the structure of the displacement — here, an error inherited from the two contact neighbours at coupling 0.7. The largest comb mean is 0.514 against a sampling band of 0.073, and the readout returns 8/13.

Errors that pass between organs

An organ's neighbours are the ones eight and thirteen places back — that is what a parastichy pair is. So a disturbance transmitted by contact is correlated at exactly the two lags the readout examines, and it does not have to be told them. Driven into a lattice with no rule in it, it returns the counted pair on eight stems out of eight.

mechanism · Noise transport
The two combs, in the proportions the rule gives them. The ratio of the second comb to the main one, for a kinematic lattice whose errors are inherited from its two contact neighbours, against how unevenly that inheritance is split. The horizontal line is where the placement rule's own stems sit, at 0.65. Weighted by distance — the coupling a d⁻³ interaction would give, which at this rise favours the 13-neighbour by 1.26 to one because the 13-hop is the shorter — the forgery sits at 1.46, well above the rule. It reaches the rule's value only at about 3 to one the other way, which is a factor of 4 against what distance supplies and in the opposite direction.

What a forgery has to know

A lattice with transported errors reproduces the comb and the pair, so one quantity is left: the two combs' relative strength. Weighted by distance the forgery puts more in the second comb than the first; the rule does the opposite. It matches only if the coupling is turned three to one towards the further neighbour, which no falloff supplies.

mechanism · Noise transport
Take away the organ eight places back, and the next one goes into the hole. The last 34 organs of a stem at a rise of 0.005, unrolled. The open circle is the organ removed — eight places before the tip. The ring at the top is where the rule puts the next organ with every organ present; the filled mark beside it is where the rule puts it with that one missing. The two are 16.4° apart, against a local spacing of 25°, and the vacancy itself is 22.7° from the undisturbed answer. Nothing else differs between the two runs: same rise, same history, same rule.

The organ that was taken away

Every observable this site has is read off an arrangement that was finished before the reading began, and earlier work here showed what that costs. So remove one primordium from a settled stem and place the next one against what is left. The rule has to answer. The rival account cannot, because in it no organ's position was ever computed from its neighbours.

mechanism · Ablation
One wrecked stem, lag by lag — golden, rise 0.005, organ 6 back. How much each lattice hop moves from organ to organ in a stem that never repaired after a single removal, over its last 120 organs. The lag-one hop is the divergence itself and it swings by 55 degrees. The lag-4 hop swings by 0.12 degrees and sits 0.01 degrees from where the undisturbed stem put it, so that one family of the original lattice is still standing organ by organ. Its multiples inherit the same steadiness and nothing else comes within a factor of twenty. The block of angles this stem repeats has a period of 4, which is the surviving lag and not a coincidence.

A period that is not a count

Eighteen wrecked stems settle into a block whose period is one of their own spiral counts, and one settles into a block of four on a lattice counted 8 and 13. The odd one is not noise. It is the case that shows what the rule is actually conserving, and it is the reason this thread is about lattice steps rather than about spirals.

lattices · Counting
A rule too long-ranged makes no pattern; every shorter one makes the same pattern. Each dot is 4 runs from a coarse start at one exponent, separated by 0.2° of placement noise. Below p ≈ 1.1 the divergence scatters by tens of degrees, which is what an arbitrary sequence gives. From p = 1.25 to p = 8 — a sixfold range — every run ends on 8/13 with the scatter between 0.75° and 1.06°.

The exponent that barely matters

An unchecked claim, repeated since the first essays, held that the repulsion's falloff exponent hardly changes the answer. It could not be tested, because the function that would have taken it never passed one down. Tested at last, it is true on a disc — by three and a half degrees across a sixteenfold range — and on a stem it decides whether there is a pattern at all.

emergence · Falloff exponent
Seven fractions with one neighbour distance and every denominator. Each member of a matched set drawn on its own stretch of the divergence axis, 0.5° either side of itself, with the nearest other rational marked. The distances are 0.3158°, 0.3158°, 0.3117°, 0.3069°, 0.3117°, 0.3077°, 0.3064° — a spread of 3.1% — while the denominators run 19, 20, 21, 23, 33, 45, 47, a factor of 2.47. That is the construction this thread needed. Every instrument for the width of a disorder dip has a free parameter set by how close the neighbour is, so a hypothesis about the neighbourhood cannot be tested by varying the neighbourhood; on this set the neighbourhood is held fixed and the arithmetic of the fraction is what varies.

Fractions with the same neighbours

Every instrument this collection has for the width of a disorder dip has a free parameter set by how close the next rational sits — which makes a hypothesis about the neighbourhood untestable with any of them. The repair is not a better instrument. It is a set of fractions whose neighbourhoods are identical and whose denominators are not, and the arithmetic supplies twenty-four of them.

tissue · Second statistic
Two combs, at a rise of 0.005. The autocorrelation of 760 divergence angles from one stem held at a rise of 0.005. The filled teeth are the lags at multiples of 8; the open teeth are the second comb, at the same spacing offset by 5. Reading the spacing off the first and the offset off the second gives the pair 8 and 13, which is what the position counter reports for the same stem — from angles alone, with no coordinate anywhere in the calculation.

The second comb

The autocorrelation of a divergence sequence has peaks at the smaller parastichy number and at every multiple of it. It also has a second set of peaks, at the same spacing, offset by the difference of the pair — so a list of angles with no coordinate in it returns both numbers rather than one.

cylinder · Sequence
Two combs, at a rise of 0.013. The autocorrelation of 760 divergence angles from one stem held at a rise of 0.013. The filled teeth are the lags at multiples of 5; the open teeth are the second comb, at the same spacing offset by 3. Reading the spacing off the first and the offset off the second gives the pair 5 and 8, which is what the position counter reports for the same stem — from angles alone, with no coordinate anywhere in the calculation.

A harmonic is a step taken twice

The spectrum contains the larger parastichy number, their sum, and echoes of the smaller one, and no ranking of peak heights separates them. What separates them is arithmetic: a harmonic is a multiple of the spacing and a family is not, and the two kinds sit in different residue classes.

cylinder · Sequence
Two shapes, two ranges, one contrast. The exponential's lattice ends at 3.63 spacings and the gaussian's at 2.25 — ranges 47% apart — and at those two ranges the contrast is 5.74 and 6.09, 6% apart. The band is what the exponent route leaves: 3.98 at p = 1, where the uncut rule makes nothing, and 7.63 at p = 1.25, where it makes a lattice.

Two shapes, one threshold

Read in the same unit, an exponential falloff and a gaussian one disagree about where the lattice ends by half. The quantity they agree on turns out to be one the earlier work measured for an unrelated reason — and it agrees with a bracket left by a sweep of a completely different parameter.

emergence · Falloff exponent
Where each kind's lattice gives way. The largest amplitude at which every run still has a lattice, and the scatter it produces there. The amplitudes are incomparable — field 0.015 (fraction of the barrier), jostle 1 (degrees of azimuth), placement 0.8 (degrees of azimuth) — and the scatters agree to 19%. The boundary belongs to the pattern rather than to the disturbance: a lattice fails at about a degree and a half of scatter, and which of three mechanisms produced it does not move where.

The boundary belongs to the pattern

Three kinds of noise, in three incommensurable units, destroy a lattice at the same place — about a degree and a half of divergence scatter. The earlier work measured that of two kinds and called it a scale rather than a constant. With a third it looks less like a coincidence and more like a property of what a lattice is.

emergence · Scatter tolerance
Four ways to count the rule's neighbourhood, and one ordering. How many organs the placement rule is looking at, as its falloff exponent is swept from 1.5 to 6, by three different definitions. Counting the organs that carry half the profile gives 33, 12, 3, 2, 1. Counting the organs that carry nine tenths gives 182, 120, 30, 8, 3. Counting the organs that carry the weighted mean lag gives 68, 45, 21, 14, 11. They disagree about the size of the neighbourhood by a factor of 10 — the widest moves by 61 times across the sweep and the narrowest by 6.1 — and every one of them falls as the exponent rises. So no choice among them changes which rule is the deeper, and no redefinition can turn a result about the depth around.

The nearest organ is not the nearest neighbour

Rank the terms of the sum the rule minimises and read off which organs the biggest ones belong to. At every falloff exponent from 1.5 to 6 the answer is the same five: lags 13, 8, 5, 21 and 26. The organ placed immediately before is not among them, and counting the neighbourhood in organs was the wrong unit.

mechanism · Noise colour
The ratio is a U across every rung, and its floor is the number that was reported. The ratio of the second comb to the main comb, on five stems at each of 15 rises spanning two rungs, against the ladder's own coordinate for where each rise sits inside its rung. Both rungs give the same shape: a floor of 0.71 and 0.79 about two thirds of the way up, climbing towards the transition at either end. The dashed line is a transported disturbance with no rule in it at 1.29, which does not vary with the rise at all — a kinematic lattice's angle sequence has no rise in it. Where the rule's curve crosses that line the two accounts are indistinguishable.

The ratio was the floor of a curve

One number was left standing between a placement rule and a transported disturbance, measured at one rise, with the explanation that the geometry there happens to favour the larger parastichy number. Swept across two rungs the number is a U — a floor of about 0.79 two thirds of the way up a rung, climbing past 2.8 as a transition approaches — and the geometry is flat exactly where the curve is steepest.

cylinder · Noise transport
How often three counters read a band's own pair, against the width of the band, on an ordinary stem. Bands of 40 to 320 organs slid up an ordinary stem grown at T = 300 over a rise from 0.05 to 0.0005, each read three ways. The counter without an index reads the pair at 65%, 66%, 87%, 88%, 88%, 88%, 89%, 100%, 100%, 92%, 54%, 23%, 8%, 2%, 1%, 0%, 0%, 0%; the counter families required to cross reads the pair at 65%, 66%, 87%, 88%, 88%, 88%, 89%, 100%, 100%, 92%, 54%, 22%, 8%, 2%, 1%, 0%, 0%, 0%; the counter with the index reads the pair at 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%, 100%. The index-free counter reads nine bands in ten or more only from 80 to 100 organs.

A counter that cannot be slid

The counter that needs no order of arrival follows each family into chains and counts them, and on an ideal lattice it agrees with the counter that does. On a stem whose rise falls it works only inside a band of widths, and outside the band it returns a pair rather than refusing: the rung below when the band is too narrow for the larger count, a pair on no rung when the band spans more than about a third of a rung of rise. The upper edge moves with the rate, so a width that is right on one stem is wrong on another, and on the fastest bijugate stem measured no width works at all.

lattices · Chain counting
What a removal costs the next organ. One mark per wrecked cut in the census: how far the first organ placed after the removal ended up from where the control put it. The rows split by which organ was taken. Removing a direct chain-neighbour of the growing tip — an organ at a multiple of one of the two counted numbers — moves the next organ by between 8.9 and 30.7 degrees. Removing anything else inside the front moves it by between 62.8 and 167.6. Nothing lands between the two groups and the ratio across the gap is 2.05, so the line is a gap rather than a threshold. Taking away a neighbour is the cheap removal, which is the opposite of what the words suggest.

Removing a neighbour costs least

Take away an organ that is a direct chain-neighbour of the growing tip and the next organ moves by under thirty-one degrees. Take away anything else inside the front and it moves by at least sixty-three. Thirty cuts, two groups, a factor of two between them and nothing in the gap.

mechanism · Ablation
The cell area a 137.508° golden head's packing is divided by, at 10 sizes. Counting every bounded cell the mean area runs from 3.607 to 38.828, the largest at 150 organs, because the cells just inside the edge of the head reach out to circumcentres far beyond it. Leaving out every cell whose polygon crosses the head's own radius, it stays between 3.1425 and 3.1634 at every size drawn, which is π, the area the square-root rule gives each organ.

Packing, measured against the interior

An earlier reading of these heads reported that no packing criterion singles out the golden angle and that three criteria give three winners. Every one of those readings was divided by a mean cell area that, on a head of 150 organs, was 38.8 where the interior's is π. Divided by the interior's own, the criteria about distance put the golden angle first of 72 angles and the criteria about cells go to rational ones.

tissue · Packing
Hold the neighbourhood and the denominator stops mattering. The equivalent width of the disorder dip — the area of the deficit divided by its own depth — for seven fractions whose nearest neighbours sit at the same distance and whose denominators run from 19 to 47. Each is measured at a head size chosen so that all of them share one scaled unit, which makes a window in scaled units the same window in degrees and the same fraction of the way to the neighbour for every member. At a window of 25 the seven widths are 38.8, 38.8, 39.0, 39.0, 38.7, 39.0, 39.0 — a spread of ×1.010 across a factor of 2.47 in denominator. The lines separate as the window widens, to ×1.147 at 200, and when they do they order by denominator rather than by crowding. So the residual this thread carried was the window: hold it and there is nothing left that belongs to the fraction.

The residual was the window

After the depth and the q over n squared scale are taken out of a disorder dip, something looked left over and looked ordered by how crowded the fraction's neighbourhood is. Measured on fractions whose neighbourhoods are identical by construction, seven widths across a factor of two and a half in denominator agree to one per cent. There is no residual; there was a comparison made at different effective windows.

wrong · Second statistic
A period of 8, and the two classes that are not with the rest. The same wrecked stem, folded on the lag it kept: one row per residue class, each drawn at the mean displacement of its own organs against the level the rest of them share. The bar through each row is the spread inside that class, and the widest of them is 0.87° — so within a class the displacement is a constant. six classes sit at the common level. The two that do not sit at 134.3° and -134.2°, equal and opposite to within 0.0 per cent, and they are neighbouring residues. The stem's own divergence is 137.44°, so an exception is one organ's step.

One level and two exceptions

Inside a wrecked stem's period most residue classes sit at one level and a couple do not. On seventeen of the thirty cuts the exceptions are exactly two, equal and opposite to within five per cent — and on all seventeen they are neighbouring residues, which was not looked for.

mechanism · Damage shape
What a lattice survives depends on the colour of the disturbance, sixfold. five stems for each of seven disturbances at each of six displacements, every stream normalised by its own measured spread so that a displacement of half a degree is half a degree in every row. A filled mark is a stem that still has a lattice — a divergence scatter under 2° — and an open one is a stem that does not. Independent errors survive to 0.5°; errors that remember the last one to 1.5°; errors inherited from the contact neighbours only to 0.25°. The number beside each row is the scatter a protractor would record where the lattice is standing, and it is the quantity that explains the table: what destroys a lattice is not how far an organ moves, but how far it moves relative to the organs it is placed against.

A disturbance the organs share

This collection has put three kinds of noise into the placement rule and found the lattice fails at about the same recorded scatter whichever kind it was. None of them asked what happens when the displacements are correlated between organs. At equal displacement per organ, a lattice survives three times as much of a disturbance the organs share — and what a protractor records is the part they do not.

emergence · Noise amplitude
What a lattice survives depends on the colour of the disturbance, sixfold. five stems for each of seven disturbances at each of six displacements, every stream normalised by its own measured spread so that a displacement of half a degree is half a degree in every row. A filled mark is a stem that still has a lattice — a divergence scatter under 2° — and an open one is a stem that does not. Independent errors survive to 0.5°; errors that remember the last one to 1.5°; errors inherited from the contact neighbours only to 0.25°. The number beside each row is the scatter a protractor would record where the lattice is standing, and it is the quantity that explains the table: what destroys a lattice is not how far an organ moves, but how far it moves relative to the organs it is placed against.

The disturbance that travels

If a lattice survives three times the displacement when the organs share it, then a disturbance passed between the organs that actually touch should be the gentlest of all — it is correlated at exactly the offsets the rule places against. It is the harshest. Half the displacement destroys what independent noise leaves standing, and the reason separates two things that had been one.

emergence · Noise colour
Removing one wall of the slot, then the other, then both. The growing tip sits in a slot between its two chain-neighbours — the organ 5 places back and the organ 8 places back. Each bar is how far the next organ placed moves when those are removed, against a control sharing the same history. Either alone is a cheap removal: 26.3° and 4.9°, both inside the 45° that separates taking a neighbour from taking anything else. Together they move it 164.1°, against 31.2° for the two effects added, so the interaction is +132.9°. The slot is not two independent walls.

Both walls of the slot

The growing tip sits between its two chain-neighbours. Removing either alone is a cheap removal on all six lattices — 2.3° to 41.7°. Removing both together throws the next organ past the expensive line on three of them, and the interaction runs from −25.8° to +132.9°.

mechanism · Both walls

The first three hundred organs

Over the counts a head shows, forty-five angles resist approximation within a per cent as well as the golden angle, and what separates them is at the centre. Grown as heads and measured there, the golden angle has the widest closest pair of all forty-six — by organs 1 and 4, the count its arithmetic names — and keeps first place only while the centre is in the reading. Its rivals stay a per cent apart from it out to a radius that tracks where their spiral counts start to add up, and every one of them is within a per cent by the 289th organ. By the largest hole it is never the best.

wrong · Hurwitz

A removal that changes nothing

On one of the six lattices, taking away both walls of the slot moves the next organ 11.953125° — which is exactly, to the last digit, what taking away the larger wall alone moves it. The smaller wall's removal contributes nothing at all when the larger one is already gone.

mechanism · Both walls

The organ that guards the second slot

An organ twelve places back is the furthest of any from where the next one goes, and removing it moves the next one by a whole divergence. The reason is that the rule's profile has two low points rather than one, the second is the slot after next, and that organ is holding it up. The comparison between what it holds up and how far behind it is decides the whole thing.

emergence · Ablation

The rung that two organs wreck

On the coarse 3/5 stem both walls of the slot heal when removed alone and wreck when removed together — and the wreck keeps no rigid hop at all. Two of the six pairs in the design end at a destination single removals almost never reach.

mechanism · Both walls

Where a handover sits

Inside every rung there is a rise at which the two contact steps change places, so that the shorter hop belongs to the other family below it. Six of the eight rungs on this ladder have one, each has exactly one, and every one of them sits in the coarse half.

cylinder · Rung interior

Two lines that cross once

The divergence at which a lattice's two contact steps are exactly equal is a curve across each rung, computable from the geometry with nothing grown. The rule's own settled divergence is a second, shallower curve, and where they cross is where the step ordering changes hands.

cylinder · Rung interior

A band that holds the angle still

Around every handover the settled divergence has a shallow floor, so a run of rises either side of it share a divergence to a twentieth of a degree while their two contact steps change places. That is a matched pair with one quantity varying, and it is the design the ablation thread had no way to state.

cylinder · Rung interior

The hop that survived

A stem that never repairs after an organ is removed settles into an exactly repeating block of angles, and the period of that block is a spiral count of the lattice it was cut from. Nobody could say why. Read the wrecked stem by lags rather than by neighbours and the answer is one line: one family of the original lattice is still standing, organ by organ, and the block is its period.

emergence · Ablation

Six lattices were not enough

The interaction between the two walls of a slot came back at −25.8° to +132.9° on six lattices, three above zero and three below, with no ordering by rise, by counted pair or by branch. A quantity that looks free on six rows is usually a quantity that has been sampled at six rows.

mechanism · Both walls

When the second wall is free

On six of thirty lattices, removing both walls of the slot costs exactly what removing the larger one alone costs — 35.9° and 35.9°, 12.0° and 12.0°, agreeing to the last digit of the grid the azimuths sit on. The smaller wall is not a wall on those rows.

mechanism · Both walls

The rung decides the sign

Twenty-four lattices where both walls of the slot are really there. Thirteen give a strongly positive interaction, at 85° to 135°; eleven give a negative or null one, at −25° to −0.5°. Nothing lies between. Every rung's lattices fall on the same side as each other.

mechanism · Both walls

A survivor has to be a neighbour

A stem that never repairs after a removal keeps exactly one lattice hop rigid, and nothing predicted which one. Sweep every offset at twelve lattices and the answer narrows sharply: at twenty-nine of thirty the surviving hop is one of the two families a counter returns, and the one exception is a step six times too long to be one.

emergence · Ablation

Not the shorter of the two

If a damaged stem keeps one contact family standing, the obvious guess is that it keeps the nearer one. Across thirty wrecked offsets that is true twelve times and false seventeen, and on one lattice the two steps differ by a quarter of a per cent — where the words shorter and longer are doing no work at all.

emergence · Ablation

The ordering was not the actor

Cut an organ out of every rise of a band where the counted pair and the settled divergence are held and the two contact steps change places, and the family left standing does not change. Fifty-five wrecked cuts on two branches, and the ordering reverses underneath every one of them.

emergence · Ablation

The organ that moved furthest

A reading that works on nine of thirty rows needs a reference organ, and the obvious repair is to measure one rather than to choose it. Measured, the disturbance turns out to have no far edge at all, so there is no organ that moved furthest in any sense the reading can use — and the generalisation that does work needs no reference organ.

emergence · Ablation

Named alongside it

The objects these essays reach for when they reach for this one.

MeasurementParastichy pairHonest limitsThe placement ruleLattice offsetDivergence angleRiseAblationLatticeClaim testingNegative resultRung

All concepts