Field

What a plant might be doing

Turing's last work was on this, and it was unpublished when he died: a ring of cells, two diffusing substances, and a spacing that selects itself. The modern account uses auxin and a pump that works uphill. Both are made to predict a number and then to produce it.
Which patterns grow on a ring of circumference 1.40. Modes 4 to 15 have positive growth rates and mode 8 is fastest. Integrating the full equations from a disordered start gives 8 peaks.

Turing's last problem

Turing's final work was on phyllotaxis and it was unpublished when he died. Its core is a ring of cells and two diffusing substances, and the thing it does is select a number of peaks — which can be predicted from the equations before anything is integrated, and then counted from what the integration produces.

The angle between the first two peaks, for six starting disorders. The same equations, the same ring, six different starting perturbations: 177°, 47°, 109°, 151°, 47°, 133°. A spiral needs the angle between successive elements to be the same one each time, and a stationary ring does not supply that.

A ring cannot make a spiral

The peaks on a Turing ring do not all appear at once — there is a first and a second. But which two lead is decided by the starting disorder, so the angle between them comes out at 177°, then 47°, then 109°, then 151°. A divergence angle is a relationship that repeats, and this one does not.

48 cells with a carrier that pumps auxin up the gradient. Each short line is one cell's polarisation — the neighbour it pumps towards, which is always the richer one. 10 peaks come out, at a contrast of 93%, from a start that was uniform to within 6%.

A pump that works uphill

The mechanism that actually has molecular support behind it does not use a diffusing inhibitor at all. Cells move auxin towards whichever neighbour already has more of it, which is the opposite of what transport is supposed to do — and it produces a spacing from a field that started uniform to within six per cent.

Which patterns grow on a ring of circumference 0.80. Modes 2 to 8 have positive growth rates and mode 4 is fastest. Integrating the full equations from a disordered start gives 4 peaks.

What a mechanism would have to show

Every model of phyllotaxis comes with the caveat that reproducing a pattern is not explaining it. That is easy to repeat and hard to make precise. Here it is made precise — a list of what an account of phyllotaxis would have to establish, with each item marked according to whether the models drawn in these essays establish it.

Three disturbances, three places to get in. The rule reads its neighbours, builds a profile of the energy at every azimuth, takes the least of it, and records a position. field noise enters at the profile; jostle noise enters at the neighbours; placement noise enters at the record. Two of the three are upstream of the choice and can change which minimum is taken; the third is downstream and never can.

The noise that arrives through the neighbours

The two kinds of noise this site had were idealisations that bracket the rule's choice. The realistic disturbance is neither: a primordium is placed exactly, and then the organ grows, so by the time the next one forms its neighbours have moved. That is a third kind, and it is invisible in every measurement a plant offers.

Three disturbances, three places to get in. The rule reads its neighbours, builds a profile of the energy at every azimuth, takes the least of it, and records a position. jostle noise enters at the neighbours; placement noise enters at the record. Two of the three are upstream of the choice and can change which minimum is taken; the third is downstream and never can.

A growing organ is part of the rule

Every model in the earlier essays places primordia on a surface and then treats the surface as furniture. But the surface grows between one placement and the next, and that growth reaches the rule through the only channel it has — where the neighbours are. What looks like a boundary condition turns out to be a term in the model.

The same lattice with no rule in it. A cylindrical lattice at a divergence of 137.826° and a rise of 0.005, built by placing node i at exactly i times the divergence and then displacing each azimuth independently by 0.5°. Its photograph is the photograph of the stem in the figure beside it and its parastichy pair is the same pair. The largest comb mean in it is 0.03 against a sampling band of 0.07, and the readout refuses.

A comb is evidence of a rule

Build the same lattice kinematically — every node at an exact multiple of the divergence, an independent error on each azimuth, no feedback anywhere — and the spectrum is empty. The photograph is identical and the parastichy pair is identical. The comb is not a property of the arrangement.

A lattice with independent errors. The autocorrelation of 759 divergence angles from a kinematic lattice at a divergence of 137.8261° and a rise of 0.005, with 0.5° of scatter on each azimuth. There is no placement rule anywhere in it: node i is put at exactly i times the divergence and then displaced. The only thing that differs between this figure and the control is the structure of the displacement — here, independent errors — that earlier work's control at 0.5° of independent scatter. The largest comb mean is 0.029 against a sampling band of 0.073, and the readout refuses.

A disturbance with a memory

That earlier work's control assumed that a plant's errors are independent from organ to organ, and nobody had tested it. Give the errors a memory — each one a fraction of the last, up to a coefficient of 0.97 — and the comb does not appear. The obvious threat to the result turns out to be empty, and the algebra says why before the measurement does.

A lattice with an error inherited from the two contact neighbours. The autocorrelation of 759 divergence angles from a kinematic lattice at a divergence of 137.8261° and a rise of 0.005, with 0.5° of scatter on each azimuth. There is no placement rule anywhere in it: node i is put at exactly i times the divergence and then displaced. The only thing that differs between this figure and the control is the structure of the displacement — here, an error inherited from the two contact neighbours at coupling 0.7. The largest comb mean is 0.514 against a sampling band of 0.073, and the readout returns 8/13.

Errors that pass between organs

An organ's neighbours are the ones eight and thirteen places back — that is what a parastichy pair is. So a disturbance transmitted by contact is correlated at exactly the two lags the readout examines, and it does not have to be told them. Driven into a lattice with no rule in it, it returns the counted pair on eight stems out of eight.

The two combs, in the proportions the rule gives them. The ratio of the second comb to the main one, for a kinematic lattice whose errors are inherited from its two contact neighbours, against how unevenly that inheritance is split. The horizontal line is where the placement rule's own stems sit, at 0.65. Weighted by distance — the coupling a d⁻³ interaction would give, which at this rise favours the 13-neighbour by 1.26 to one because the 13-hop is the shorter — the forgery sits at 1.46, well above the rule. It reaches the rule's value only at about 3 to one the other way, which is a factor of 4 against what distance supplies and in the opposite direction.

What a forgery has to know

A lattice with transported errors reproduces the comb and the pair, so one quantity is left: the two combs' relative strength. Weighted by distance the forgery puts more in the second comb than the first; the rule does the opposite. It matches only if the coupling is turned three to one towards the further neighbour, which no falloff supplies.

Take away the organ eight places back, and the next one goes into the hole. The last 34 organs of a stem at a rise of 0.005, unrolled. The open circle is the organ removed — eight places before the tip. The ring at the top is where the rule puts the next organ with every organ present; the filled mark beside it is where the rule puts it with that one missing. The two are 16.4° apart, against a local spacing of 25°, and the vacancy itself is 22.7° from the undisturbed answer. Nothing else differs between the two runs: same rise, same history, same rule.

The organ that was taken away

Every observable this site has is read off an arrangement that was finished before the reading began, and earlier work here showed what that costs. So remove one primordium from a settled stem and place the next one against what is left. The rule has to answer. The rival account cannot, because in it no organ's position was ever computed from its neighbours.

The ratio follows the disturbance, not the rule. The ratio of the second comb to the main comb on stems grown by the placement rule and jostled by seven different disturbances, all at 0.25° of displacement per organ and all on the same rule. Independent errors and errors with a memory return 0.76–0.81, which is the value this site measured for the rule. A periodicity at the smaller parastichy number takes it down to 0.45; errors inherited from the contact neighbours take it up to 1.09, most of the way to the 1.24 a transported disturbance gives with no rule in it at all. So the quantity separates arrangements by how their errors are related, not by whether anything computed the positions.

The ratio was never about the rule

The comb has already been retracted here as evidence that a plant computes its pattern, and one quantity was exempted from the retraction: the ratio of the two combs, which a placement rule and a transported disturbance divide differently. Drive seven disturbances through the same rule and the ratio spans 0.45 to 1.09. The exemption does not hold, and the angle sequence has nothing left.

What the model settles on, against how fast the meristem grows. A broad golden branch, a transition, and then the two-whorl regime at exactly half a turn. 16 of 40 converged settings land within 4° of the golden angle; 17 land more than 20° away.

Two-ranked, by two different routes

The rule produces a two-ranked stem at a coarse rise, where 180° is the only thing available, and that has been in the bifurcation diagram since the beginning. It also produces one at a fine rise, at a rise whose own answer is the golden angle, if a single organ is removed. The diagram cannot show the second, and the reason it cannot is how it is drawn.

Two combs, at a rise of 0.005. The autocorrelation of 760 divergence angles from one stem held at a rise of 0.005. The filled teeth are the lags at multiples of 8; the open teeth are the second comb, at the same spacing offset by 5. Reading the spacing off the first and the offset off the second gives the pair 8 and 13, which is what the position counter reports for the same stem — from angles alone, with no coordinate anywhere in the calculation.

A disturbance that is not passed on

The disturbance that forges every observable measured here does two things at once — it correlates an organ's error with its contact neighbours', and it hands that error on to be handed on again. Every result about it has been unable to say which half did the work. This is the control that takes the second half away and keeps the first.

The damage is the sharing; the forgery is the history. Three disturbances of the same size, measured four ways. The two left columns are stems grown by the placement rule and jostled at 0.25° per organ: a disturbance shared between the contact neighbours scatters the lattice by 0.71° against white noise's 0.57°, and one inherited from them — the same sharing, passed on again at every organ — by 0.97°. The two right columns are kinematic lattices with no rule in them at all, where the whole question is what a disturbance can manufacture. The inherited one returns the pair on 8 seeds of 8 with a main comb of 0.205 against a band of 0.073; the shared one, at the same coupling and the same scatter, returns it on 1 and makes a comb of 0.099, which is the band. So sharing an error with the organs you touch does the damage, and only passing it on and on forges the evidence.

The forgery needs a history

A disturbance passed between touching organs manufactures the comb, the second comb and the parastichy pair on an arrangement with no rule in it — which is why the comb stopped being evidence. Give the organs the same correlation with no accumulation in it and the forgery collapses: one seed in eight returns a pair, and the comb is the noise floor.

The wander is in the disturbance and not in what a plant lets you measure. Each disturbance measured twice, in the same statistic. On the left, the variance of the block means of the disturbance's own deviates, over blocks of 100, as a multiple of what independent draws would give; on the right, the same quantity for the divergence sequence those deviates produce, over blocks of 128. The left column is what this site measured when it proposed a slow wander as a second observable. The right column is what a botanist would have: a divergence is the difference of two organs' errors, and differencing is exactly the operation that removes power at low frequencies. The disturbance inherited between touching organs goes from ×49.1 — the largest here — to 0.83, which is what independent errors give. The one with a memory in time keeps most of its own.

A difference forgets a drift

This collection proposed a second observable and priced it as free: if a plant's errors are inherited between touching organs, the divergence sequence should carry a slow wander as well as a comb. The disturbance with the largest wander of any built here leaves none at all in the sequence, because a divergence is a difference and differencing is what removes a drift.

Through the rule, the drift survives and the inheritance still does not. How much of a divergence sequence's variance survives being averaged over blocks, on stems the rule grew. The vertical quantity is B² times the variance of the block means divided by the variance of the sequence, which is one at every block size for independent errors — the arithmetic is normalised for a differenced stream, since a divergence is the difference of two organs' errors. A line that climbs is a sequence with power at frequencies below one per block. The disturbances that remember the last error climb to 46 at a block of 64. The ones inherited between touching organs do not climb at all — 1.51 and 1.90 at the same block — although their own deviates carry ×— and ×— an independent stream's variance in exactly this statistic. What they do instead is dig a hole: at block sizes of 8 and 13, which are the offsets they couple at, the statistic falls to 0.06 and 0.12.

What the rule does to a drift

A placement rule was supposed to leave no slow wander in a divergence sequence, because its errors are corrections rather than inheritances. Driven by a disturbance that drifts, it leaves a larger one than a lattice with no rule in it at all — while cutting the per-organ scatter by more than half. The rule removes what is relative between neighbours, and a drift is not.

The loop bound is not the neighbourhood. The wander a placement rule leaves in its divergences, against how many recently placed organs the rule sums over, at four correlation lengths of the disturbance driving it. The loop runs from 15 organs to 85 and nothing moves: the largest change along any line is smaller than the change between random seeds at one setting. That is the shape a parameter has when it is not binding, and it is the same shape a robust result has, which is why the sweep is drawn with the seed spread rather than reported as a number.

The window was not the neighbourhood

A placement rule corrects what is relative between neighbours and passes what moves them all together, so how much of a slow disturbance gets through should depend on how deep the neighbourhood is. The obvious knob is how many organs the rule sums over. Swept across a factor of six, it changes nothing at all — and a parameter that is not binding produces exactly the flat sweep a robust result produces.

A deeper rule passes more of a drift, not less. The wander left in a stem's divergences, against how many organs its disturbance stays correlated over, for rules whose neighbourhoods run from 3 organs to 182. The prediction under test said a rule should pass a drift once the drift outlasts its neighbourhood, so the shallow rules should be the leaky ones and each line should turn where its own depth is crossed. Every line rises smoothly and the deepest rule is the highest of them at every correlation length — 82 against 22 at the longest drift. There is no crossover anywhere in the sweep.

The drift goes the other way

A rule that corrects what its neighbourhood shares should let through any disturbance slower than its own reach, and should suppress anything faster — a crossover, tracking the depth. Swept over a neighbourhood that changes by a factor of sixty, there is no crossover anywhere, and the deep rule passes nearly four times as much as the shallow one. The prediction is not weakly supported; it is backwards.

The wander climbs because its denominator falls. Three quantities across the same sweep of the rule's depth, each divided by its own value for the shallowest rule so that they share an axis. The wander rises by a factor of 3.7 as the neighbourhood goes from 3 organs to 182. The scatter between neighbouring divergences falls by 1.57, because that is the part of a disturbance a placement rule corrects and a deeper rule corrects it better. What is left — how many degrees of slow drift actually reach the divergences — changes by 1.20, from 1.52 to 1.82 degrees. The rule barely filters a drift at any depth.

What the ratio was hiding

The statistic that says a rule sharpens a drift rises by a factor of nearly four across a sweep of the rule's depth. Undo the normalisation and ask instead how many degrees of drift actually reach the divergences, and the answer changes by a fifth. Nearly all of the effect was in the denominator, and the denominator is the thing the rule is good at.

A sixfold neighbourhood, and nothing to dilute. The prediction was that a rule with fewer neighbours would convert a jostle into divergence scatter more efficiently. Across a sixfold widening the ratio sits between 0.97 and 1.00, and the one point that differs is the narrowest, at 0.82 — smaller, where the prediction wanted larger. The row underneath is why: past four spacings the rule builds the identical lattice, internode for internode, so there is no neighbourhood left to widen.

The neighbourhood was already settled

The earlier work explained a small difference between two kinds of noise by saying a jostle is diluted among some thirty neighbours. Sweep the neighbourhood sixfold and the difference does not move — because past four spacings the rule builds the identical lattice, internode for internode. There was nothing to dilute.

Four ways to count the rule's neighbourhood, and one ordering. How many organs the placement rule is looking at, as its falloff exponent is swept from 1.5 to 6, by four different definitions. Counting the organs that carry half the profile gives 33, 12, 3, 2, 1. Counting the organs that carry nine tenths gives 182, 120, 30, 8, 3. Counting the organs that carry ninety-nine hundredths gives 284, 263, 149, 54, 9. Counting the organs that carry the weighted mean lag gives 68, 45, 21, 14, 11. They disagree about the size of the neighbourhood by a factor of 10 — the widest moves by 61 times across the sweep and the narrowest by 6.1 — and every one of them falls as the exponent rises. So no choice among them changes which rule is the deeper, and no redefinition can turn a result about the depth around.

Four ways to count a neighbourhood

How deep does the placement rule look? Counting the organs that carry nine tenths of its profile gives 182 down to 3 as the falloff steepens. Counting the ones that carry half gives 33 down to 1. The weighted mean lag gives 68 down to 11. The four disagree by an order of magnitude about the size and agree exactly about the order.

Four ways to count the rule's neighbourhood, and one ordering. How many organs the placement rule is looking at, as its falloff exponent is swept from 1.5 to 6, by three different definitions. Counting the organs that carry half the profile gives 33, 12, 3, 2, 1. Counting the organs that carry nine tenths gives 182, 120, 30, 8, 3. Counting the organs that carry the weighted mean lag gives 68, 45, 21, 14, 11. They disagree about the size of the neighbourhood by a factor of 10 — the widest moves by 61 times across the sweep and the narrowest by 6.1 — and every one of them falls as the exponent rises. So no choice among them changes which rule is the deeper, and no redefinition can turn a result about the depth around.

The nearest organ is not the nearest neighbour

Rank the terms of the sum the rule minimises and read off which organs the biggest ones belong to. At every falloff exponent from 1.5 to 6 the answer is the same five: lags 13, 8, 5, 21 and 26. The organ placed immediately before is not among them, and counting the neighbourhood in organs was the wrong unit.

Which rule passes more drift — exponent 1.5 against exponent 6. The same disturbance is given to two placement rules, one with a falloff exponent of 1.5 and one of 6, and the bar counts how many of six seeds let more of it through under the deeper rule. Above the line means the deeper rule passed more. Reading left to right the disturbance is given a longer memory, from white noise to a correlation length of 99 organs. The two change hands: the shallower rule passes more up to a correlation length of 1.4 organs and the deeper one from 2.8 organs onward. The comparison is made seed by seed rather than between two averages, because the spread between seeds at one setting is larger than the difference being measured.

The corner that does not move

Read as degrees of drift getting through rather than as a ratio, and compared seed by seed, the deep and shallow rules change hands. The share that goes to the deeper rule climbs from twenty-three per cent under white noise to ninety-four at a correlation length of a hundred organs — and the crossing sits at two or three organs whether the two rules differ by a factor of four or sixty-one.

What a removal costs the next organ. One mark per wrecked cut in the census: how far the first organ placed after the removal ended up from where the control put it. The rows split by which organ was taken. Removing a direct chain-neighbour of the growing tip — an organ at a multiple of one of the two counted numbers — moves the next organ by between 8.9 and 30.7 degrees. Removing anything else inside the front moves it by between 62.8 and 167.6. Nothing lands between the two groups and the ratio across the gap is 2.05, so the line is a gap rather than a threshold. Taking away a neighbour is the cheap removal, which is the opposite of what the words suggest.

Removing a neighbour costs least

Take away an organ that is a direct chain-neighbour of the growing tip and the next organ moves by under thirty-one degrees. Take away anything else inside the front and it moves by at least sixty-three. Thirty cuts, two groups, a factor of two between them and nothing in the gap.

A period of 5, and the two classes that are not with the rest. The same wrecked stem, folded on the lag it kept: one row per residue class, each drawn at the mean displacement of its own organs against the level the rest of them share. The bar through each row is the spread inside that class, and the widest of them is 10.28° — so within a class the displacement is a constant. three classes sit at the common level. The two that do not sit at 143.0° and -147.6°, equal and opposite to within 3.2 per cent, and they are neighbouring residues. The stem's own divergence is 137.97°, so an exception is one organ's step.

The damage has a period

Every wrecked stem in the census has had two numbers read out of its displacement profile and the profile itself read out of none of them. Folded on the lag the stem kept, twenty-five of the thirty are constant inside each residue class to between 0.12° and 6.09°.

A period of 8, and the two classes that are not with the rest. The same wrecked stem, folded on the lag it kept: one row per residue class, each drawn at the mean displacement of its own organs against the level the rest of them share. The bar through each row is the spread inside that class, and the widest of them is 0.87° — so within a class the displacement is a constant. six classes sit at the common level. The two that do not sit at 134.3° and -134.2°, equal and opposite to within 0.0 per cent, and they are neighbouring residues. The stem's own divergence is 137.44°, so an exception is one organ's step.

One level and two exceptions

Inside a wrecked stem's period most residue classes sit at one level and a couple do not. On seventeen of the thirty cuts the exceptions are exactly two, equal and opposite to within five per cent — and on all seventeen they are neighbouring residues, which was not looked for.

The exceptional pair, measured in divergences. One row per wrecked cut whose profile has exactly one pair of exceptional classes, drawn at the size of that pair divided by the stem's own settled divergence. Every row sits between 0.882 and 1.076, so the two chains that came apart moved by one organ's step rather than by two or by half of one. The residual is not scatter: rows are grouped by the lag the stem kept, and every lag sits wholly above the line or wholly below it. Why a surviving 5 or 7 overshoots and a surviving 4 or 8 falls short is not answered here.

A step of one organ

The balanced pair inside a wrecked stem's period measures 88.0° to 147.2° against divergences of 99.1° to 138.0° — one organ's step, to within twelve per cent on every row. The residual is not scatter: every stem keeping a 5 or a 7 overshoots and every stem keeping a 4 or an 8 falls short.

How constant the displacement is inside one residue class. One row per wrecked cut in the census, drawn at the widest spread found inside any one residue class when the profile is folded on the lag that stem kept. 25 of 30 rows sit between 0.12 and 6.09 degrees, which on a quantity whose between-class differences run past a hundred and fifty degrees is a constant. The five that do not sit from 10.3° up. There is nothing in between, so the line drawn at 10° could have been drawn anywhere in a wide interval.

The plateau was a prediction

The search for a reference organ found that the largest displacement above a hole is a plateau rather than a peak, and reported it as a failure. A profile constant on each of k residue classes has exactly k levels, so its maximum is attained by a whole class — a ninth to a quarter of every window, forever.

How far above the hole the damage becomes a pattern. One row per wrecked cut, drawn at the first organ from which every residue class stays at its own level for the rest of the run. On the 25 rows that reach it at all, it runs from 7 to 303 organs above the removed one; five rows never reach it inside the 300 organs each run is continued for. Below that point the stem is still moving, and the displacement of the first organ after the cut — the quantity that tells a cheap removal from an expensive one — is measured there. Above it, nothing changes again.

Two regimes above a hole

Below the repeating pattern there is a transient, and the boundary between them is measurable: the first organ from which every class stays at its own level runs from 7 to 303 organs above the hole on twenty-five of thirty cuts, and five never reach it inside the run.

Removing one wall of the slot, then the other, then both. The growing tip sits in a slot between its two chain-neighbours — the organ 5 places back and the organ 8 places back. Each bar is how far the next organ placed moves when those are removed, against a control sharing the same history. Either alone is a cheap removal: 26.3° and 4.9°, both inside the 45° that separates taking a neighbour from taking anything else. Together they move it 164.1°, against 31.2° for the two effects added, so the interaction is +132.9°. The slot is not two independent walls.

Both walls of the slot

The growing tip sits between its two chain-neighbours. Removing either alone is a cheap removal on all six lattices — 2.3° to 41.7°. Removing both together throws the next organ past the expensive line on three of them, and the interaction runs from −25.8° to +132.9°.

Removing one wall of the slot, then the other, then both. The growing tip sits in a slot between its two chain-neighbours — the organ 5 places back and the organ 8 places back. Each bar is how far the next organ placed moves when those are removed, against a control sharing the same history. Either alone is a cheap removal: 25.8° and 12.0°, both inside the 45° that separates taking a neighbour from taking anything else. Together they move it 12.0°, against 37.7° for the two effects added, so the interaction is -25.8°. The slot is not two independent walls.

A removal that changes nothing

On one of the six lattices, taking away both walls of the slot moves the next organ 11.953125° — which is exactly, to the last digit, what taking away the larger wall alone moves it. The smaller wall's removal contributes nothing at all when the larger one is already gone.

Which hops survive one wall, the other, and both. One row per lattice. The last three columns are the lags whose hop the cut stem still holds, unchanged from a control that shares its history — the measurement that identifies what a wrecked stem has become. Removing a single wall always leaves something standing, which is what every single-organ cut in this collection does. Removing both leaves nothing at all on two of six lattices, including the coarse rung that no single removal can wreck. A stem that keeps no rigid hop is not a wrecked lattice with a slip in it; it is a stem that is no longer a lattice.

The rung that two organs wreck

On the coarse 3/5 stem both walls of the slot heal when removed alone and wreck when removed together — and the wreck keeps no rigid hop at all. Two of the six pairs in the design end at a destination single removals almost never reach.

Which chain the backward exception sits on, over the census. Chains are numbered from the removed organ, so chain 0 is the chain the hole was on and chain 2 is two organs along it. The exchange is at the hole's own chain on 10 of the 17 rows that carry one, against 2.8 rows for a chain drawn at random from each row's own period. That is far more often than anywhere else and it is not every row, so the position is a tendency rather than a rule — and the file says so rather than rounding it up.

Which chains changed places

A wrecked stem's displacement profile is a set of levels, one per chain, with two of them out of line — equal and opposite, on neighbouring chains. Nothing said which two. They are the hole's own chain and the one below it, on ten of the seventeen cuts that carry a pair.

A period of 8, with the hole's own chain at the top. Each mark is one residue class of the displacement profile, placed round a ring at its own residue, with the chain the removed organ sat on at the top. The radius is how far that class sits from the level the rest of them share. six of the eight classes sit together at the middle ring; two do not, and on this row they are one pair, equal and opposite to within a twentieth. The forward one is chain 7 and the backward one is chain 0, one residue above it, which is the order every row of the census puts them in.

One way round, seventeen times

The two chains that change places in a wrecked stem are adjacent, which is symmetric and says nothing about direction. Label them by lag from the hole and the one displaced forwards is always the lower of the two — on every row of the census, without an exception.

Four accounts of one angle, scored on the same 17 rows. Each bar is how far an account of the exchanged pair's size sits from the measurement, as a share of the control's divergence, averaged over the census. A step of the wrecked stem's own divergence is the obvious candidate and the worst of the four. The surviving family's own step is not the unit at all: that angle is a fifth of the exchange. A step of the control's divergence is close, and correcting it by a fifth of the surviving hop takes the worst row from 11.8 per cent to 4.0.

Four accounts of one angle

The exchanged pair in a wrecked stem is about one divergence step, and about is doing twelve per cent of work. Four candidate units were written down and scored on the same seventeen rows: the cut stem's own step, the surviving family's step, the control's step, and the control's corrected.

What the exchange misses one divergence step by, against the surviving hop. Each mark is one wrecked cut. Across: the angle from an organ to the one its surviving lag places above it, on the control. Down: how much the exchanged pair's size falls short of one divergence step. The relationship is a straight line through the origin with a slope of about 0.2, and its sign is right on every row of the census — the four surviving lags give hops on both sides of zero and the residual follows each of them. The honest limit is the four: the hop hardly moves inside a lag, so the line rests on 11 distinct positions and not on 17.

A fifth of the hop

The exchanged pair misses one divergence step by up to twelve per cent, and the miss is not scatter: every row keeping a lag of 5 or 7 overshoots and every row keeping a 4 or an 8 falls short. Subtract a fifth of the surviving hop's own angle and the worst row is four per cent.

What removing both walls does that removing each does not. One row per lattice, drawn at the difference between how far the next organ moves when both walls of the slot are removed and how far the two removals move it separately added together. Zero would mean the walls act independently. five of six lattices are more than 10° from it, three above and two below — so the pair is reliably not the sum of its parts and is not reliably larger than it either. On four of them the pair throws the next organ past the 45° that separates a cheap removal from an expensive one, which neither wall alone comes near.

Six lattices were not enough

The interaction between the two walls of a slot came back at −25.8° to +132.9° on six lattices, three above zero and three below, with no ordering by rise, by counted pair or by branch. A quantity that looks free on six rows is usually a quantity that has been sampled at six rows.

Where taking the second wall as well changes nothing. Each row is one lattice, with three marks: how far the next organ moves when the smaller wall alone is removed, when the larger alone is removed, and when both are. On these lattices the third mark sits on the second, to within two steps of the azimuth grid. The smaller wall is free — taking it away as well changes nothing — and on a row like that the interaction is minus the smaller wall's own cost by construction, which is arithmetic and not a measurement. Three of them are the whole of one rung and the others are the fine ends of two more.

When the second wall is free

On six of thirty lattices, removing both walls of the slot costs exactly what removing the larger one alone costs — 35.9° and 35.9°, 12.0° and 12.0°, agreeing to the last digit of the grid the azimuths sit on. The smaller wall is not a wall on those rows.

Five accounts of the sign, on the 24 lattices that are measurements. Each bar is how many of the lattices an account puts on the right side of zero. The six rows where the second wall is free are left out, because their value is minus the first wall's cost by construction and any rule scores whatever it happens to say about them. Position inside the rung, the rise and the branch all fail. The larger counted number sorts 22 of the 24, and the misses are one rung's worth of rows rather than a scatter.

The rung decides the sign

Twenty-four lattices where both walls of the slot are really there. Thirteen give a strongly positive interaction, at 85° to 135°; eleven give a negative or null one, at −25° to −0.5°. Nothing lies between. Every rung's lattices fall on the same side as each other.

The interaction across each rung, coarse end to fine end. One line per rung, drawn against where in the rung each lattice sits — nought at the coarse end, one at the fine end, measured in the logarithm of the rise. The lines are flat. Inside a rung the interaction moves by 3.3 to 13.6 degrees, against a spread of 240 degrees across the ladder, and it falls from the coarse end to the fine one on 6 of the 7 rungs. Position inside a rung was the candidate this design was built to test and it is not what decides the sign.

The exception was already labelled

The larger counted number sorts twenty-two of twenty-four lattices by the sign of their slot interaction. Both misses are on the Lucas 3/4 rung — the one rung a different measurement had already singled out, for reasons with nothing to do with this one.

Where the pattern starts, measured at two run lengths. One row per wrecked cut. The small mark is the onset a 300-organ run reports and the ring is what a 600-organ run reports; a row with only one mark reports an onset at only one length. 19 of the 24 rows that report both move by more than twenty organs, and the largest move is from 135 to 435. The range the thread has been quoting, 7 to 303 organs, becomes 28 to 473.

Twice the run

Five wrecked cuts never reached a pattern inside three hundred organs and one reached it at three hundred and three, which is a number asking to be checked. Run every cut in the census twice as far and three of the thirty change their answer.

Where the pattern starts, measured at two run lengths. One row per wrecked cut. The small mark is the onset a 300-organ run reports and the ring is what a 600-organ run reports; a row with only one mark reports an onset at only one length. 19 of the 24 rows that report both move by more than twenty organs, and the largest move is from 135 to 435. The range the thread has been quoting, 7 to 303 organs, becomes 28 to 473.

An onset at the end of the run

One cut reported that its pattern began 299 organs into a 300-organ run. Given twice the room it reports no pattern at all. The reading was the run stopping, not the disturbance ending, and the definition guarantees one at the last organ of every run.

The 3 cuts the two run lengths disagree about. Each block is one wrecked cut, with its widest within-class spread drawn at both run lengths and the 10 degrees that separates periodic from not marked by the rule. Two of these become periodic when the run is doubled, at spreads falling from about seventy degrees to about eight. One goes the other way, from six degrees to a hundred and seventy — and that one is the row an entirely separate reading of the same census independently reports as its worst fit.

Three rows change sides

Twenty-five of thirty wrecked cuts have a periodic displacement profile over three hundred organs and twenty-six do over six hundred. The count barely moves and the membership does: two rows join, one leaves, and the gap the threshold sits in narrows from 1.69 to 1.27.

How steady each class is, over 300 organs and over 600. Each mark is one wrecked cut, placed across at the widest spread found inside any one of its residue classes over the shorter run and up at the same reading over the longer one. A mark on the diagonal is a row the two lengths agree about. The rules are the 10 degrees that separates a profile called periodic from one that is not: three rows fall in different quadrants at the two lengths, two of them becoming periodic and one ceasing to be. The gap between the two groups narrows from 1.69 times to 1.27.

A window nobody aligned

Every reading this thread takes of a wrecked stem is taken inside a window, and there are three of them: a run of three hundred organs, a window of a hundred and twenty at its top, and a search window of fifteen to thirty-nine. None was aligned to anything, and one of them turned out to decide its own answers.

Three chains, each moved a third of a turn, on g008 cut at offset 6. The eight chains of a stem that kept a lag of 8, set round the circle, with the three that are displaced marked and their displacements written on. Each is about a third of a turn the same way round, and they sum to -359.6 degrees — one whole turn to within half a degree, on an azimuth grid whose step is a quarter of a degree. A rearrangement that closes by going once round sums to a turn, not to nothing, which is why the test it was set had the wrong number in it.

A cycle sums to a whole turn

The test proposed for whether three displaced chains are a three-cycle was that their displacements sum to zero. Three chains rotating into one another's places each move about a third of a turn the same way round, and a third of a turn three times is a whole turn — which the unfolded test calls the worst row in the census.

Each cut's class spread at three reading windows. The worst spread within a residue class, measured over the last 60, 120 and 180 organs of the same runs. A quantity that is steady with noise on it would give a flat line; a quantity drifting steadily along the run gives a line through the origin. Nearly every row gives the second, so the classes this thread calls steady are sliding slowly and the number that says how much depends on how much of the run is read. The dashed guide is exact proportionality.

A spread that grows with its window

A spread over a sample of a steady quantity does not depend on how big the sample is. These spreads triple when the window triples, on nearly every row of the census, which means the classes this thread calls steady are sliding — slowly, and invisibly at any single window.

How far each rearrangement is from closing, against the lag it kept. One bar per excluded cut, its length the distance from the displacements summing to nothing, ordered smallest first, and each labelled with the lag the cut left standing. The seven shortest bars are the cuts that kept a lag of 7 or 8 and the six longest are the cuts that kept 4 or 5, with a gap of 6.3 degrees between the two groups and no row in it. The rule sorts every one of them.

The lag decides whether it closes

Seven of the thirteen excluded rows have displacements that cancel and six do not. Every row that closes kept a lag of seven or eight and every row that does not kept four or five, thirteen times out of thirteen — and then a lattice nobody had cut broke it.

The 13 cuts the exchange sets aside, chain by chain. One row per wrecked cut with no balanced pair, one cell per chain of the lag it kept, and the chains displaced away from the common level filled. The lag is printed at the left and the sum of the displacements at the right, folded into half a turn either way. seven of the 13 sum to within 12 degrees of nothing and six do not, and every row that closes kept a lag of 7 or 8 while every row that does not kept 4 or 5. The rows are stacked by that lag.

The rows nobody added up

Seventeen of the census's thirty wrecked cuts come back as one balanced pair of displaced chains, and every claim about the exchange is quantified over those seventeen. The other thirteen were set aside as having three or more exceptions and never looked at again. They are one addition each.

Which cuts count as periodic, at each reading window. One row per wrecked cut and one column per window. A filled cell is a cut whose worst class spread is under the ten-degree line and is therefore called periodic. At 60 organs 28 of the 30 cuts are, at 120 organs 25, and at 180 organs 25. three rows change side, all of them losing their periodicity as the window widens, and they are marked.

The window nobody moved

Three instrument settings sit between the ablation census and every statement it makes. Two have been varied and both decided answers. The third is a hundred and twenty organs at the top of a run, it has never been moved, and moving it changes the verdict on three rows.

Where each window begins, against where the pattern begins. The run is drawn left to right, one bar per cut. The pale stretch is the disturbance still healing, up to the organ from which the profile holds its levels for the rest of the run; the dark stretch is the pattern repeating. The marks are the organ each of the three windows starts at, since a window is the last N organs of a run. On these rows the widest window starts inside the pale stretch, so it is reading the healing and calling it the pattern — which is why widening the window takes them off the periodic list.

Three rows a window moves

Three of the census's thirty cuts are periodic when a hundred and twenty organs are read and not when a hundred and eighty are. Their spreads do not grow in proportion to the window, they grow by twenty and forty times, and the reason is that a window is the tail of a run.

The eight rows where the two levels disagree. Each wrecked cut's residue classes drawn as points on a circle of angle, one panel per row. A tick is one class's mean displacement. The solid radius is the densest cluster of them, which is the level this file computes; the pale radius is the median of the class means, which is the level everything in the ablation thread has been measured against. On every one of these rows the median stands on a class that is on its own, and on all 28 rows not drawn the two levels agree to within the exception tolerance of 10 degrees.

A median that is an exception

Every number in the ablation thread is measured against the value most of a wrecked stem's chains sit at, taken as a median so that a few exceptions cannot move it. On eight of thirty-six rows the median stands on a chain sitting by itself.

No majority and no balanced pair, over the whole census. One row per wrecked cut. The bar is the share of that row's residue classes sitting at the level they agree on, and the vertical rule is a half — a bar reaching past it has a majority and a median is safe there. The mark at the right says whether the exchange keeps that row. 15 rows are on both lists of 16 and 16, and the two part on g008/6, which is excluded and has a majority of 5 of 8, and l013/4, whose lag is 4 so that a balanced pair leaves two classes each way and a majority is arithmetically unavailable.

No majority and no pair

Whether a wrecked stem's chains agree on a level and whether its exceptions form a balanced pair are computed from different things. Sixteen rows have neither, sixteen have neither, and fifteen rows are in both — and the two they part on are the two most interesting rows in the census.

g005 at offset 6, read against two levels. One wrecked cut's four residue classes, drawn twice. On the left the level is the median of the four class means, which falls on the class at -5.5 degrees sitting on its own; three classes are then exceptions and the row is set aside as having no balanced pair, summing to -173.6 degrees. On the right the level is the pair of classes that agree, at -138.6 degrees; two classes are then exceptions, they are adjacent and equal and opposite at 133.1 and 134.2 degrees, and the row is an exchange. The exchange table goes from 20 rows to 21.

The twenty-first row

Recomputing the level moves one row out of the set the exchange sets aside and into the exchange itself. Its hop is four times larger than any the correction was fitted over, and the correction fails on it in the one way it had never failed.

How far each set-aside row's rearrangement is from closing, at both levels. Every row the exchange sets aside, placed by how far its displacements sum from nothing after folding. A mark below the line kept a lag of seven or more and one above kept four or five, which is the rule that sorts them. At the median level the rule is wrong on one row of 16, at -22.6 degrees, and the two populations are 6.3 degrees apart. At the recomputed level it is right on every one of 15 — one row having left the set entirely — and the gap is 3.2 degrees, with the nearest open row at 12.1 against a threshold of 12.

The rule comes back

A rule sorting the awkward rows was right fifteen times in sixteen, with the one failure blamed on a statistic. Recomputing the statistic makes it fifteen of fifteen — and halves the gap the line is drawn in, which is the price.

Predicted 29 of 63 settle; 17 do. The 63 wrecked runs of the slot design regrown to 1200 organs and put to the settling table's own criterion, unchanged in every tolerance. Rows are whether the 300-organ endpoint sat within 1 degree of a destination; columns are whether the regrown run settles. The prediction written down before the sweep was that the 29 agreeing runs would settle: 12 of them do and 17 refuse, and the whole set settles 17 rather than 29. Agreement carries information without being a rule — 41.4 per cent of the agreeing runs settle against 14.7 per cent of the others, an odds ratio of 4.09 and a phi coefficient of 0.300 — so an agreeing run is still likelier to refuse than to settle.

A destination or a refusal

Sixty-three wrecked runs were regrown to twelve hundred organs and put to the settling table's own criterion, unchanged in every tolerance. The prediction written down before the sweep said twenty-nine would settle; seventeen do, and the prediction is wrong on its own side of the table as well as in its total.

The endpoint over four times the run: 60 of 63 do not move at all. One cell per wrecked run, dark where the divergence reported as its endpoint at 300 organs is the same divergence at 1200, warm where it moved. It moves on three of 63 runs, by a median of 0.0000 degrees and a worst of 0.0039. That worst is 128 times smaller than the 0.5 degrees at which two settled values are counted as one destination, and the movement has no sign — 1 up, 2 down. The short reading is precise about where a run finishes; it is silent about whether the run stays there.

Like with like

Seventeen wrecked runs settle when regrown to four times their length, and every one of them settles at the divergence the short run already reported. The endpoint moves by four thousandths of a degree at worst, which makes the short reading precise about where and silent about whether.

The same verdict at every length from 300 organs to 1200. The settling criterion applied to the first 300, 400, 600, 800, 1000, 1200 organs of each of the 63 regrown runs. It accepts 17 at every one of them and refuses 46 at every one, and not one run changes sides. A truncated twelve-hundred-organ run is the shorter run organ for organ, so this is a comparison of lengths rather than of growths. The slowest arrival in the whole set is 138 organs, so 140 is the measured requirement and 300 carries a factor of 2.1. The column drawn dark is 300 organs, where the verdict is 17 settled and 46 refused.

What a run length was hiding

The settling criterion returns an identical verdict on all sixty-three wrecked runs at every length from three hundred organs to twelve hundred, so run length explains nothing. What the sweep does find is that an endpoint is a mean over an orbit, and thirty-four refusers never come within a degree of their own reported endpoint.

Six boundaries on three basins, five kinds between them, and three that are edges. Each basin's stretch of starting angle, with both its boundaries located to ± 0.125° by sweeping ten degrees at a quarter of a degree. The pale bar behind each is the interval the forty-angle table could bracket it in, three to five degrees at a time. The widest basin, at rise 0.03 and exponent 2, is a fringe at 124.375° and a puncture at 179.625°. The same-rise basin, at rise 0.03 and exponent 3, is a sliver at 124.875° and a fade at 167.625°. The narrow basin, at rise 0.02 and exponent 3, is a wall at 144.875° and a fade at 161.625°. Only 3 of the 6 are edges in the sense of a side: the widest basin's upper boundary is a hole 0.75° wide centred on 180°, with the same destination beyond it, so that basin runs out of basin at the reflection point rather than reaching an edge. The interval the forty-angle table bracketed each basin in is drawn behind it, from the sweep at 1200 organs a run.

A basin with no upper edge

The widest basin in the settling table had a width bracketed between 47.5 degrees and about 57, and closing a bracket means sampling near an edge rather than everywhere. Three basins cut at a quarter of a degree located all six of their boundaries, and the widest turned out to run out of basin at 180 degrees rather than reach an edge on that side at all.

Two instruments at six boundaries: the clock cannot tell a wall from a fade and the tail spread separates them by a factor of 327. The clock is the ratio of the slowest settling in the last degree inside a basin to the middle of the rest of its window. It rises towards every boundary and diverges at none: the one clean wall rises by 1.313× and the two clean fades by 1.00× and 1.57×, so the wall sits between them and no threshold on a clock separates the two kinds. The tail spread is how far a run's own last two hundred organs wander, and every run has one whether it settles or not. Past the wall it is 0.101°, as steady as the basin just left; past the two fades it is 33.042° and 39.712°. Pooled over all 574 runs everything that settles spreads by 0–0.442° and everything that does not by 30.828–39.712°. The two instruments are drawn side by side on one row per boundary, from the sweep at 1200 organs a run.

A wall or a fade

A basin's border is either a change of destination or a stretch where the angles stop settling at all, and nothing here could tell the two apart. Two instruments were pointed at the question: the settling clock, which looked obviously right and fails, and the tail spread, which was already being computed on every run and had never been read.

1.75° of starting angle between two basins that reaches neither, and a 3° void beside it. 20.25 degrees of starting angle swept unbroken at 0.25°, from inside the basin at 101.5° to inside the widest basin at 139.3°, one cell per sampled angle. The last angle reaching 101.5° is 114.5° and the first reaching 139.3° is 116.25°; the 1.75° between them holds 6 sampled angles and 0 of them settle anywhere at all. The widest run of angles reaching nothing is 3° wide, at 118–120.75°, and a detached island of 139.3° sits between the two. Starting angle is left over. The stretch that reaches neither basin is bracketed above the strip, from the sweep at 1200 organs a run.

The angles left over

A stem started anywhere on the circle was assumed to end up in one basin or another, so that the settled destinations divided the starting angles between them. Twenty and a quarter degrees swept without a hole at a quarter of a degree find 1.75 degrees between two basins that reaches neither of them and nothing else, and a three-degree void beside it.

41.75° in the middle of the widest basin, sampled only at the table's own 3.12–4.38°. The widest basin from its located lower boundary at 124.375° to the reflection point at 180°, with the two ten-degree windows swept at 0.25° shaded and the stretch between them left open. Nothing has looked inside that stretch more finely than the forty-angle table's own 3.12–4.38° spacing. The narrowest feature this sweep found anywhere is the 0.75° wedge inside same-rise, and a feature that size falls between the table's angles 79% of the time — 75% under an even 3° sampling. So an unmeasured sliver or puncture could sit anywhere in the middle of this basin and nothing here would have seen it. The two swept windows are shaded and the stretch between them is left open, from the sweep at 1200 organs a run.

What a quarter degree cannot see

Six boundaries were located to an eighth of a degree, three basins were named and one width was quoted, and every one of those readings has the same floor under it. The sweep's grid is one step of the grid the stems are placed on, so nothing here bounds a basin narrower than half a degree — and the widest basin's own middle was never swept at all.

Ten of the 24 orderings four exponents admit, from 15 readings of the wall. Every arrangement of the four falloff exponents is a cell, and a cell is filled when some level of some sampling puts the four walls in that order. 15 readings of runs that are shared cell for cell give ten of the 24, with the most common occurring three times. The marked cells are the orderings read at a half, the only level any round of this collection has published, and each of them occurs once. Exponent 5 is ranked first in 8 of the 15 readings and each of the others in two or three.

The level was doing the ordering

Four falloff exponents have been put in order by the rise at which half their runs stop settling, and a half is the only level that order has ever been read at. Read at nine levels the same runs give ten different orderings of the same four numbers, and the one comparison in the whole study that resolves runs the other way.

Every bracket on the wall at a half, before the refinement and after. One pair of bars per falloff exponent: above, the rises consistent with that exponent's crossing on forty starting angles and the 5 published rises; below, the same on eighty angles and 9 rises. Two of the four have an open end before — at a half that is the coarse end, where no published rise has a share confidently above the level — and none is open after, the widest closing at 1.57 times. The four walls sit inside a factor of 1.111 of one another, against 1.624 on the reading that could not locate them, so they came closer together rather than further apart.

Four walls closer than they looked

Two of the four falloff exponents had a wall with no upper end at all, and the other two were located to factors of two and a half and nearly four. Nine rises at eighty starting angles close every bracket — and the four walls turn out to sit inside a factor of 1.111 of one another, which is narrower than the narrowest bracket.

The bracket at each of the four corners of the refinement, at a half. The geometric mean bracket at each corner, and under it how much of the joint narrowing each half of the refinement accounts for. It is taken over the two exponents whose bracket is closed at every corner, with the two halves of the refinement crossed against each other. The rule across the bars is 1.54, the narrowest bracket the published rise list can express at all. On forty starting angles and the published rises it is 3.06 times with two of the four brackets closed; doubling the angles alone gives 1.96, halving the rise spacing alone gives 1.46, and both together give 1.44. In the logarithm, where the two effects add, the rises account for 98.8 per cent of the joint narrowing and the angles for 59.3 per cent — summing to 158.1 per cent, which is what overlapping causes look like and not two independent factors of two.

Two refinements that do not multiply

The design that located the wall did two things at once — doubled the starting angles and halved the rise spacing — and the arithmetic behind it assumed each would buy about a factor of two. The finer rises did ninety-nine per cent of the narrowing and the doubled angles added under one, because a bracket's ends are rises and no error bar can move them.

The settling share against the rise at exponent 5, on the refined rises. One line, over 80 starting angles at exponent 5. The rise runs coarse on the left and the level the wall is read at is the dashed rule. Between 0.02 and 0.013 the refined list holds two rises the published one does not, and at exponent 5 the share reaches 0.725 at 0.01732 against 0.550 at the ends of that stretch — 3.13 standard errors above the higher end. It turns in all three readings independently, which is what separates a feature of the curve from a bump in a sample. The ringed point is that maximum.

A maximum in the gap

Four falloff exponents have refused to separate on every quantity this thread has read off them, and the wall that was supposed to tell them apart cannot. Two of the four carry a maximum in the settling share at a rise the published list stepped straight over, and it is there in both halves of the sampling independently.

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