What a plant might be doing
Turing's last problem
Turing's final work was on phyllotaxis and it was unpublished when he died. Its core is a ring of cells and two diffusing substances, and the thing it does is select a number of peaks — which can be predicted from the equations before anything is integrated, and then counted from what the integration produces.
A ring cannot make a spiral
The peaks on a Turing ring do not all appear at once — there is a first and a second. But which two lead is decided by the starting disorder, so the angle between them comes out at 177°, then 47°, then 109°, then 151°. A divergence angle is a relationship that repeats, and this one does not.
A pump that works uphill
The mechanism that actually has molecular support behind it does not use a diffusing inhibitor at all. Cells move auxin towards whichever neighbour already has more of it, which is the opposite of what transport is supposed to do — and it produces a spacing from a field that started uniform to within six per cent.
What a mechanism would have to show
Every model of phyllotaxis comes with the caveat that reproducing a pattern is not explaining it. That is easy to repeat and hard to make precise. Here it is made precise — a list of what an account of phyllotaxis would have to establish, with each item marked according to whether the models drawn in these essays establish it.
The noise that arrives through the neighbours
The two kinds of noise this site had were idealisations that bracket the rule's choice. The realistic disturbance is neither: a primordium is placed exactly, and then the organ grows, so by the time the next one forms its neighbours have moved. That is a third kind, and it is invisible in every measurement a plant offers.
A growing organ is part of the rule
Every model in the earlier essays places primordia on a surface and then treats the surface as furniture. But the surface grows between one placement and the next, and that growth reaches the rule through the only channel it has — where the neighbours are. What looks like a boundary condition turns out to be a term in the model.
A comb is evidence of a rule
Build the same lattice kinematically — every node at an exact multiple of the divergence, an independent error on each azimuth, no feedback anywhere — and the spectrum is empty. The photograph is identical and the parastichy pair is identical. The comb is not a property of the arrangement.
A disturbance with a memory
That earlier work's control assumed that a plant's errors are independent from organ to organ, and nobody had tested it. Give the errors a memory — each one a fraction of the last, up to a coefficient of 0.97 — and the comb does not appear. The obvious threat to the result turns out to be empty, and the algebra says why before the measurement does.
Errors that pass between organs
An organ's neighbours are the ones eight and thirteen places back — that is what a parastichy pair is. So a disturbance transmitted by contact is correlated at exactly the two lags the readout examines, and it does not have to be told them. Driven into a lattice with no rule in it, it returns the counted pair on eight stems out of eight.
What a forgery has to know
A lattice with transported errors reproduces the comb and the pair, so one quantity is left: the two combs' relative strength. Weighted by distance the forgery puts more in the second comb than the first; the rule does the opposite. It matches only if the coupling is turned three to one towards the further neighbour, which no falloff supplies.
The organ that was taken away
Every observable this site has is read off an arrangement that was finished before the reading began, and earlier work here showed what that costs. So remove one primordium from a settled stem and place the next one against what is left. The rule has to answer. The rival account cannot, because in it no organ's position was ever computed from its neighbours.
The ratio was never about the rule
The comb has already been retracted here as evidence that a plant computes its pattern, and one quantity was exempted from the retraction: the ratio of the two combs, which a placement rule and a transported disturbance divide differently. Drive seven disturbances through the same rule and the ratio spans 0.45 to 1.09. The exemption does not hold, and the angle sequence has nothing left.
Two-ranked, by two different routes
The rule produces a two-ranked stem at a coarse rise, where 180° is the only thing available, and that has been in the bifurcation diagram since the beginning. It also produces one at a fine rise, at a rise whose own answer is the golden angle, if a single organ is removed. The diagram cannot show the second, and the reason it cannot is how it is drawn.
A disturbance that is not passed on
The disturbance that forges every observable measured here does two things at once — it correlates an organ's error with its contact neighbours', and it hands that error on to be handed on again. Every result about it has been unable to say which half did the work. This is the control that takes the second half away and keeps the first.
The forgery needs a history
A disturbance passed between touching organs manufactures the comb, the second comb and the parastichy pair on an arrangement with no rule in it — which is why the comb stopped being evidence. Give the organs the same correlation with no accumulation in it and the forgery collapses: one seed in eight returns a pair, and the comb is the noise floor.
A difference forgets a drift
This collection proposed a second observable and priced it as free: if a plant's errors are inherited between touching organs, the divergence sequence should carry a slow wander as well as a comb. The disturbance with the largest wander of any built here leaves none at all in the sequence, because a divergence is a difference and differencing is what removes a drift.
What the rule does to a drift
A placement rule was supposed to leave no slow wander in a divergence sequence, because its errors are corrections rather than inheritances. Driven by a disturbance that drifts, it leaves a larger one than a lattice with no rule in it at all — while cutting the per-organ scatter by more than half. The rule removes what is relative between neighbours, and a drift is not.
The window was not the neighbourhood
A placement rule corrects what is relative between neighbours and passes what moves them all together, so how much of a slow disturbance gets through should depend on how deep the neighbourhood is. The obvious knob is how many organs the rule sums over. Swept across a factor of six, it changes nothing at all — and a parameter that is not binding produces exactly the flat sweep a robust result produces.
The drift goes the other way
A rule that corrects what its neighbourhood shares should let through any disturbance slower than its own reach, and should suppress anything faster — a crossover, tracking the depth. Swept over a neighbourhood that changes by a factor of sixty, there is no crossover anywhere, and the deep rule passes nearly four times as much as the shallow one. The prediction is not weakly supported; it is backwards.
What the ratio was hiding
The statistic that says a rule sharpens a drift rises by a factor of nearly four across a sweep of the rule's depth. Undo the normalisation and ask instead how many degrees of drift actually reach the divergences, and the answer changes by a fifth. Nearly all of the effect was in the denominator, and the denominator is the thing the rule is good at.
The neighbourhood was already settled
The earlier work explained a small difference between two kinds of noise by saying a jostle is diluted among some thirty neighbours. Sweep the neighbourhood sixfold and the difference does not move — because past four spacings the rule builds the identical lattice, internode for internode. There was nothing to dilute.
Four ways to count a neighbourhood
How deep does the placement rule look? Counting the organs that carry nine tenths of its profile gives 182 down to 3 as the falloff steepens. Counting the ones that carry half gives 33 down to 1. The weighted mean lag gives 68 down to 11. The four disagree by an order of magnitude about the size and agree exactly about the order.
The nearest organ is not the nearest neighbour
Rank the terms of the sum the rule minimises and read off which organs the biggest ones belong to. At every falloff exponent from 1.5 to 6 the answer is the same five: lags 13, 8, 5, 21 and 26. The organ placed immediately before is not among them, and counting the neighbourhood in organs was the wrong unit.
The corner that does not move
Read as degrees of drift getting through rather than as a ratio, and compared seed by seed, the deep and shallow rules change hands. The share that goes to the deeper rule climbs from twenty-three per cent under white noise to ninety-four at a correlation length of a hundred organs — and the crossing sits at two or three organs whether the two rules differ by a factor of four or sixty-one.
Removing a neighbour costs least
Take away an organ that is a direct chain-neighbour of the growing tip and the next organ moves by under thirty-one degrees. Take away anything else inside the front and it moves by at least sixty-three. Thirty cuts, two groups, a factor of two between them and nothing in the gap.
The damage has a period
Every wrecked stem in the census has had two numbers read out of its displacement profile and the profile itself read out of none of them. Folded on the lag the stem kept, twenty-five of the thirty are constant inside each residue class to between 0.12° and 6.09°.
One level and two exceptions
Inside a wrecked stem's period most residue classes sit at one level and a couple do not. On seventeen of the thirty cuts the exceptions are exactly two, equal and opposite to within five per cent — and on all seventeen they are neighbouring residues, which was not looked for.
A step of one organ
The balanced pair inside a wrecked stem's period measures 88.0° to 147.2° against divergences of 99.1° to 138.0° — one organ's step, to within twelve per cent on every row. The residual is not scatter: every stem keeping a 5 or a 7 overshoots and every stem keeping a 4 or an 8 falls short.
The plateau was a prediction
The search for a reference organ found that the largest displacement above a hole is a plateau rather than a peak, and reported it as a failure. A profile constant on each of k residue classes has exactly k levels, so its maximum is attained by a whole class — a ninth to a quarter of every window, forever.
Two regimes above a hole
Below the repeating pattern there is a transient, and the boundary between them is measurable: the first organ from which every class stays at its own level runs from 7 to 303 organs above the hole on twenty-five of thirty cuts, and five never reach it inside the run.
Both walls of the slot
The growing tip sits between its two chain-neighbours. Removing either alone is a cheap removal on all six lattices — 2.3° to 41.7°. Removing both together throws the next organ past the expensive line on three of them, and the interaction runs from −25.8° to +132.9°.
A removal that changes nothing
On one of the six lattices, taking away both walls of the slot moves the next organ 11.953125° — which is exactly, to the last digit, what taking away the larger wall alone moves it. The smaller wall's removal contributes nothing at all when the larger one is already gone.
The rung that two organs wreck
On the coarse 3/5 stem both walls of the slot heal when removed alone and wreck when removed together — and the wreck keeps no rigid hop at all. Two of the six pairs in the design end at a destination single removals almost never reach.
Which chains changed places
A wrecked stem's displacement profile is a set of levels, one per chain, with two of them out of line — equal and opposite, on neighbouring chains. Nothing said which two. They are the hole's own chain and the one below it, on ten of the seventeen cuts that carry a pair.
One way round, seventeen times
The two chains that change places in a wrecked stem are adjacent, which is symmetric and says nothing about direction. Label them by lag from the hole and the one displaced forwards is always the lower of the two — on every row of the census, without an exception.
Four accounts of one angle
The exchanged pair in a wrecked stem is about one divergence step, and about is doing twelve per cent of work. Four candidate units were written down and scored on the same seventeen rows: the cut stem's own step, the surviving family's step, the control's step, and the control's corrected.
A fifth of the hop
The exchanged pair misses one divergence step by up to twelve per cent, and the miss is not scatter: every row keeping a lag of 5 or 7 overshoots and every row keeping a 4 or an 8 falls short. Subtract a fifth of the surviving hop's own angle and the worst row is four per cent.
Six lattices were not enough
The interaction between the two walls of a slot came back at −25.8° to +132.9° on six lattices, three above zero and three below, with no ordering by rise, by counted pair or by branch. A quantity that looks free on six rows is usually a quantity that has been sampled at six rows.
When the second wall is free
On six of thirty lattices, removing both walls of the slot costs exactly what removing the larger one alone costs — 35.9° and 35.9°, 12.0° and 12.0°, agreeing to the last digit of the grid the azimuths sit on. The smaller wall is not a wall on those rows.
The rung decides the sign
Twenty-four lattices where both walls of the slot are really there. Thirteen give a strongly positive interaction, at 85° to 135°; eleven give a negative or null one, at −25° to −0.5°. Nothing lies between. Every rung's lattices fall on the same side as each other.
The exception was already labelled
The larger counted number sorts twenty-two of twenty-four lattices by the sign of their slot interaction. Both misses are on the Lucas 3/4 rung — the one rung a different measurement had already singled out, for reasons with nothing to do with this one.
Twice the run
Five wrecked cuts never reached a pattern inside three hundred organs and one reached it at three hundred and three, which is a number asking to be checked. Run every cut in the census twice as far and three of the thirty change their answer.
An onset at the end of the run
One cut reported that its pattern began 299 organs into a 300-organ run. Given twice the room it reports no pattern at all. The reading was the run stopping, not the disturbance ending, and the definition guarantees one at the last organ of every run.
Three rows change sides
Twenty-five of thirty wrecked cuts have a periodic displacement profile over three hundred organs and twenty-six do over six hundred. The count barely moves and the membership does: two rows join, one leaves, and the gap the threshold sits in narrows from 1.69 to 1.27.
A window nobody aligned
Every reading this thread takes of a wrecked stem is taken inside a window, and there are three of them: a run of three hundred organs, a window of a hundred and twenty at its top, and a search window of fifteen to thirty-nine. None was aligned to anything, and one of them turned out to decide its own answers.
A cycle sums to a whole turn
The test proposed for whether three displaced chains are a three-cycle was that their displacements sum to zero. Three chains rotating into one another's places each move about a third of a turn the same way round, and a third of a turn three times is a whole turn — which the unfolded test calls the worst row in the census.
A spread that grows with its window
A spread over a sample of a steady quantity does not depend on how big the sample is. These spreads triple when the window triples, on nearly every row of the census, which means the classes this thread calls steady are sliding — slowly, and invisibly at any single window.
The lag decides whether it closes
Seven of the thirteen excluded rows have displacements that cancel and six do not. Every row that closes kept a lag of seven or eight and every row that does not kept four or five, thirteen times out of thirteen — and then a lattice nobody had cut broke it.
The rows nobody added up
Seventeen of the census's thirty wrecked cuts come back as one balanced pair of displaced chains, and every claim about the exchange is quantified over those seventeen. The other thirteen were set aside as having three or more exceptions and never looked at again. They are one addition each.
The window nobody moved
Three instrument settings sit between the ablation census and every statement it makes. Two have been varied and both decided answers. The third is a hundred and twenty organs at the top of a run, it has never been moved, and moving it changes the verdict on three rows.
Three rows a window moves
Three of the census's thirty cuts are periodic when a hundred and twenty organs are read and not when a hundred and eighty are. Their spreads do not grow in proportion to the window, they grow by twenty and forty times, and the reason is that a window is the tail of a run.
A median that is an exception
Every number in the ablation thread is measured against the value most of a wrecked stem's chains sit at, taken as a median so that a few exceptions cannot move it. On eight of thirty-six rows the median stands on a chain sitting by itself.
No majority and no pair
Whether a wrecked stem's chains agree on a level and whether its exceptions form a balanced pair are computed from different things. Sixteen rows have neither, sixteen have neither, and fifteen rows are in both — and the two they part on are the two most interesting rows in the census.
The twenty-first row
Recomputing the level moves one row out of the set the exchange sets aside and into the exchange itself. Its hop is four times larger than any the correction was fitted over, and the correction fails on it in the one way it had never failed.
The rule comes back
A rule sorting the awkward rows was right fifteen times in sixteen, with the one failure blamed on a statistic. Recomputing the statistic makes it fifteen of fifteen — and halves the gap the line is drawn in, which is the price.
A destination or a refusal
Sixty-three wrecked runs were regrown to twelve hundred organs and put to the settling table's own criterion, unchanged in every tolerance. The prediction written down before the sweep said twenty-nine would settle; seventeen do, and the prediction is wrong on its own side of the table as well as in its total.
Like with like
Seventeen wrecked runs settle when regrown to four times their length, and every one of them settles at the divergence the short run already reported. The endpoint moves by four thousandths of a degree at worst, which makes the short reading precise about where and silent about whether.
What a run length was hiding
The settling criterion returns an identical verdict on all sixty-three wrecked runs at every length from three hundred organs to twelve hundred, so run length explains nothing. What the sweep does find is that an endpoint is a mean over an orbit, and thirty-four refusers never come within a degree of their own reported endpoint.
A basin with no upper edge
The widest basin in the settling table had a width bracketed between 47.5 degrees and about 57, and closing a bracket means sampling near an edge rather than everywhere. Three basins cut at a quarter of a degree located all six of their boundaries, and the widest turned out to run out of basin at 180 degrees rather than reach an edge on that side at all.
A wall or a fade
A basin's border is either a change of destination or a stretch where the angles stop settling at all, and nothing here could tell the two apart. Two instruments were pointed at the question: the settling clock, which looked obviously right and fails, and the tail spread, which was already being computed on every run and had never been read.
The angles left over
A stem started anywhere on the circle was assumed to end up in one basin or another, so that the settled destinations divided the starting angles between them. Twenty and a quarter degrees swept without a hole at a quarter of a degree find 1.75 degrees between two basins that reaches neither of them and nothing else, and a three-degree void beside it.
What a quarter degree cannot see
Six boundaries were located to an eighth of a degree, three basins were named and one width was quoted, and every one of those readings has the same floor under it. The sweep's grid is one step of the grid the stems are placed on, so nothing here bounds a basin narrower than half a degree — and the widest basin's own middle was never swept at all.
The level was doing the ordering
Four falloff exponents have been put in order by the rise at which half their runs stop settling, and a half is the only level that order has ever been read at. Read at nine levels the same runs give ten different orderings of the same four numbers, and the one comparison in the whole study that resolves runs the other way.
Four walls closer than they looked
Two of the four falloff exponents had a wall with no upper end at all, and the other two were located to factors of two and a half and nearly four. Nine rises at eighty starting angles close every bracket — and the four walls turn out to sit inside a factor of 1.111 of one another, which is narrower than the narrowest bracket.
Two refinements that do not multiply
The design that located the wall did two things at once — doubled the starting angles and halved the rise spacing — and the arithmetic behind it assumed each would buy about a factor of two. The finer rises did ninety-nine per cent of the narrowing and the doubled angles added under one, because a bracket's ends are rises and no error bar can move them.
A maximum in the gap
Four falloff exponents have refused to separate on every quantity this thread has read off them, and the wall that was supposed to tell them apart cannot. Two of the four carry a maximum in the settling share at a rise the published list stepped straight over, and it is there in both halves of the sampling independently.